41Ranunculus repens, Creeping buttercup, Crowfoot, Goldcup, Kingcup.Ubiquitous in <strong>Ashdown</strong> <strong>Forest</strong>. It is <strong>of</strong>ten found in <strong>the</strong> richer, disturbed places, and like R.acris avoids <strong>the</strong> acidic soils. The grassland plants <strong>of</strong>ten differ from <strong>the</strong> finer, smaller plantsfound in wet flushes in woodland around <strong>the</strong> <strong>Forest</strong>, and it is known to be a very variablespecies, even within populations (Coles 1977),Found throughout Europe and Asia,Ranunculus bulbosus. Bulbous buttercup, Crowfoot, Goldcup, Kingcup.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980), with many additional records from <strong>the</strong> Weald in Briggs(1990).Locally frequent on road verges and in unimproved meadows around <strong>the</strong> <strong>Forest</strong>, floweringjust before creeping buttercup and a slightly different colour; if in doubt <strong>the</strong> reflexed sepalsshould be distinctive (but see R. sardous). Also rarely found on rides and grassy areasamongst hea<strong>the</strong>r as near Stone Cottage, 1995, AK.It occurs throughout Europe and locally in western Asia and North Africa.4 5Ranunculus sardous (R. hirsutus). Hairy buttercup.Fisher's Gate (Wolley-Dod 1937). Tetrads 33W and 53B (Hall 1980). Tetrads 43H and 43W(Briggs 1990) and locally frequent on <strong>the</strong> moist <strong>Ashdown</strong> Sands.This species is quite characteristic <strong>of</strong> pastures on <strong>the</strong> clays and arable land in <strong>the</strong> MedwayValley, where it may be locally abundant. There were large populations near Friars Gate in3 -++---------'+- 1987/88, and in 1991 one horse-grazed field was a sea <strong>of</strong> yellow so it is presumably not eatenlike <strong>the</strong> o<strong>the</strong>r buttercups. Our records are mainly from <strong>the</strong> edge <strong>of</strong> <strong>the</strong> Medway valley, with oneplant on a verge near Gills Lap. It is ra<strong>the</strong>r like R. bulbosus with its reflexed sepals, but itflowers for a longer period, has more yellowy-green leaves and has a few tubercles on <strong>the</strong>4 5fruits (absent in R. bulbosusl.Most frequent in west and central Europe, also in western Asia and North Africa.Ranunculus sceleratus. Celery-leaved buttercup.Pond by road, Tile Barn Farm (477.335), 1993, MM (<strong>the</strong> 33W tetrad record in Hall 1980 maybe from outside our area).This species is very characteristic <strong>of</strong> damp mud on <strong>the</strong> edges <strong>of</strong> ponds, or sometimestrampled marshes in fields, and is beautifully adapted to its temporary habitat. It is a summer3 annual or less <strong>of</strong>ten a winter annual, and can even produce two generations in a field season.It can form a persistent seed bank, and germination is enhanced by fluctuating temperaturesand exposure to sunlight. The plants can flower and die within two months, some producing4 5 up to 56,000 seeds. The achenes adhere well to animals and birds, and can also float forseveral days to aid dispersal to nearby habitats (Grime, Hodgson & Hunt 1988).Widespread in Britain, Europe, Asia, North Africa and North America.*Ranunculus lingua. Greater spearwort.--;--===-----,--, Around pond at spring at Chelwood Vachery (432.297), 1994, TR; one plant introduced witho<strong>the</strong>r aquatics to a pond on <strong>the</strong> golf course, <strong>Forest</strong> Row (437.342), found on 14 August 1995and destroyed when pond cleared out <strong>the</strong> following day, TR & PW; introduced by A. Morriss toPippingford Park under pontoon bridge (449.297) and at small pond (453.316), 1995, TR.3 -++-__....----'--1-This plant <strong>of</strong> water margins and pond edges is currently spreading in Britain (Rich &Woodruff 1996), presumably from plants thrown out <strong>of</strong> gardens. It is an uncommon native <strong>of</strong>fens. It also occurs in Europe (rarer in <strong>the</strong> south) and in Asia.4 5
42Ranunculus f1ammula subsp. f1ammula. Lesser spearwort.Wet places in many habitats, common (Hall 1980).Quite common on wet rides, around ponds and in flushes in meadows and woodland, and<strong>of</strong>ten where <strong>the</strong>re are some nutrients, but not in eutrophic places. Exasperatingly absent froma number <strong>of</strong> <strong>the</strong> higher, drier squares such as Gills Lap to Kings Standing (we have looked).Able to tolerate ferrous iron and <strong>of</strong>ten in such flushes.Common in Britain and Europe, but decreasing in England (Rich & Woodruff 1996). Italso occurs in temperate Asia and <strong>the</strong> Azores.Ranunculus ficaria. Lesser celandine.One <strong>of</strong> <strong>the</strong> earliest species to flower, and locally abundant on roadsides, in woodland, on stream banks, etc., <strong>the</strong> brightyellow flowers opening widely in sunshine. The mild winter and wet <strong>of</strong> 1994/5 was beneficial for its growth and <strong>the</strong>rewere excellent displays in spring 1995. We have searched <strong>the</strong> <strong>Forest</strong> for it specially in March and April, and most <strong>of</strong> <strong>the</strong>squares which lack records probably lack <strong>the</strong> plant, although we have not had access to all land at <strong>the</strong> right time <strong>of</strong> year.The leaves go yellow and are harder to find by end <strong>of</strong> May, and are usually gone by July.Widespread in Sussex, Britain and Europe. Sell (1994) distinguished four subspecies in Britain, <strong>of</strong> which three occur inour <strong>Flora</strong> area. They are best distinguished by flower size and <strong>the</strong> presence or absence <strong>of</strong> axillary bulbi Is (best seen afterflowering as <strong>the</strong>y can be very small and overlooked before). We have attempted to map <strong>the</strong>m wherever practical.i) Subsp. ficaria has small petals usually to 15 mm on <strong>the</strong> <strong>Forest</strong> (to 20 mm elsewhere) and lacks bulbi Is. It is widespreadaround <strong>the</strong> <strong>Forest</strong> and edges, <strong>of</strong>ten as scattered plants. Judging from <strong>the</strong> records it is about as common as <strong>the</strong> followingsubspecies.ii) Subsp. bulbi/ifer also has small petals usually to 11 mm and has bulbi Is in <strong>the</strong> leafaxils from which it regenerates, and<strong>of</strong>ten forms large patches in woodlands, gardens and waste ground (e.g. behind <strong>the</strong> telephone box at Wych Cross,419.319). Most <strong>of</strong> <strong>the</strong> records are from around <strong>the</strong> edges <strong>of</strong> <strong>the</strong> <strong>Forest</strong> and in urban areas.iii) *Subsp. chrysocephalus has petals to 23 mm (flowers 47 mm diameter) and lacks bulbils. It was found on dumped soilnear <strong>the</strong> Half Moon (498.334), 1995, TR, <strong>the</strong> first record for Sussex, and it looked very big by comparison with <strong>the</strong> o<strong>the</strong>rtwo subspecies. It appears to be native in <strong>the</strong> east Mediterranean region, and could be widespread 'In <strong>British</strong> gardens,A double-flowered garden form (taxon unknown) was found dumped and established by School Lane, Nutley(443.280), 1995, TR & JK.Ranunculus ficaria Subsp. (!caria Subsp. bulbilifer Subsp. chrysocephalus3 -+~-.....----'L-J-j- 3 -++-----'-+-4 54 5Ranunculus hederaceus. Ivy-leaved crowfoot.Marden's Hill, E. D. Morgan (Wolley-Dod 1937). Tetrads 42T and 43K (Hall 1980).Alas, we have not refound this plant on <strong>the</strong> <strong>Forest</strong>, and it seems to be quite rare now in Sussex. It is a stronglywestern oceanic species occurring from sou<strong>the</strong>rn Sweden to Spain, and is endemic to Europe.It can be distinguished from <strong>the</strong> following species by <strong>the</strong> leaves which are widest above <strong>the</strong> base and petals usuallyless than 4 mm (leaves widest at <strong>the</strong> base and petals usually more than 5 mm in R. omiophyllus). It usually grows insimilar wet muddy places, <strong>of</strong>ten where <strong>the</strong>y are disturbed and open.Ranunculus omiophyllus (R. lenormandii). Round-leaved crowfoot.Nutley 1851, J. Weaver; <strong>Ashdown</strong> <strong>Forest</strong>, C. E. Salmon; near Crowborough, E. Ellman(Wolley-Dod 1937). Tetrads 42J, 42N, 43A, 43L and 43R (Hall 1980).Shaded stream north-west <strong>of</strong> Cri pps Manor {398.3211, 1995, AK; Chelwood Gate area(41.29), 1995, MM; mud by stream near <strong>Ashdown</strong> Park (428.3191, 1993, TR & EL; three] -+i.;.--------'-+_ sites along <strong>the</strong> main stream Ii) shaded mud, Braberry Hatch (428.296), 1993, TR & EL, (ii)4 5Mill Wood (437.288), 1994, ER, RN et al. and (iii) alder wood Newbridge {455.3231, 1993, TR& PD; marshy pond in Toll Lane woodland (461.262), 1993, TR & NM where it has beenknown since 1983 AFRR; Duddleswell, shady stream by footbridge (467.283), 1995, RN &ER; <strong>Forest</strong> Row, recently cleared pond {436.3421, 1995, TR & PS.Our records are generally from mud in shaded habitats and possibly for good reason. Cook (1966) notes that inBritain its distribution follows <strong>the</strong> 8 cm August rainfall isohyet, and it is absent from much <strong>of</strong> <strong>the</strong> drier central and easternEngland; our shaded sites may be those which remain cool and damp enough for it in <strong>the</strong> summer. Elsewhere it occurs on
- Page 1 and 2: Flora of Ashdown Forest,-,;I
- Page 3 and 4: Copyright Tim Rich, Pat Donovan, Pa
- Page 5: ACKNOWLEDGEMENTSWe would like to th
- Page 8 and 9: THE HABITATS OF ASH DOWN FOREST3His
- Page 12 and 13: Near the bottom of all the non~wood
- Page 14 and 15: Conopodium majusCrataegus laevigata
- Page 16 and 17: Fincham (1995) published a small ph
- Page 18 and 19: 13ANAL VS IS OF THE FLORA DATAIn th
- Page 20 and 21: 15The number of different visits to
- Page 22 and 23: 17Hedgerows Disturbed/waste ground
- Page 24 and 25: 19The frequencies with which specie
- Page 26 and 27: 21(r~0.434, pO.l).-0.059,•300t350
- Page 28 and 29: 23A second type of comparison has b
- Page 30 and 31: 25Similarly, environmental variable
- Page 32 and 33: SPECIES ACCOUNTS27Lat'1n names foll
- Page 34 and 35: 29EQUISETACEAEEquisetum f/uviatile.
- Page 36 and 37: One plant on a vertical, north-faci
- Page 38 and 39: Viola pa/ustris in an open alder co
- Page 40 and 41: 35DRYOPTERIDACEAEPolystichum aculea
- Page 42 and 43: 37BLECHNACEAEBlechnum spicant. Hard
- Page 44 and 45: 39AngiospermsLAURACEAE* Laurus nobi
- Page 48 and 49: wet mud, small streams, ditches and
- Page 50 and 51: 45*Meconopsis cambrica. Welsh poppy
- Page 52 and 53: 47Ulmusx hollandica f?U. glabra x m
- Page 54 and 55: 49FAGACEAEFagus sylvatica. Beech.Fo
- Page 56 and 57: A special large oak at Chuck Hatch,
- Page 58 and 59: Cory/us avel/ana. Hazel.Recorded in
- Page 60 and 61: plants in the autumn; we have not r
- Page 62 and 63: Sagina procumbens. Procumbent pearl
- Page 64 and 65: 59*Saponaria officinalis. Soapwort,
- Page 66 and 67: 61Dolygonum arenastrum. Equal-leave
- Page 68 and 69: 63Rumex conglomeratus. Clustered do
- Page 70 and 71: 65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
- Page 84 and 85: 79aphanus raphanistrum. Wild radish
- Page 86 and 87: ica tetralix. Cross~leaved heath.As
- Page 88 and 89: centre of the Forest on alluvial so
- Page 90 and 91: It is predominantly a plant which o
- Page 92 and 93: 87SAXIFRAGACEAE[*Darmera pe/tata. I
- Page 94 and 95: 89insipid in flavour IHaskel1 1960)
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91ection Rubus. Subsection Hiemales
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93Rubus pr%ngatus.Nutley (45.27), 1
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95bus vestitus.Newbridge, 1969, B.
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97Rubus largificus.3 -+t--------'+_
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Potentilla anserina. Silverweed.Com
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101Geum urbanum. Wood avens.Primari
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103osa x pseudorusticana fR. arvens
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105Prunus spinosa. Blackthorn, Wint
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orbus aucuparia. Rowan, Mountain as
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109r*Crataegus persimilis. Broad-le
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111icia tetrasperma. Smooth tare.Fo
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113Trifolium rep ens. White clover"
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115Genista tinctoria. Dyer's greenw
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*Genista anglica. Petty whin, Needl
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Myriophyllum alterniflorum. Alterna
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121Epilobium montanum x obscurum.Fo
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habitats.It lives up to the name of
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125,UPHORBIACEAEVlercurialis perenn
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Although one of Britain's tiniest p
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129*Oxalis articulata. Pink-sorrel.
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*Erodium cicutarium. Common stork's
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Widespread in Europe from the shore
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* Sison amomum. Stone parsley.Tetra
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137Centaurium pu/chellum. Lesser ce
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139CONVOLVULACEAEConvolvulus arvens
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141*Nymphoides pe/tata. Fringed wat
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Myosotis secunda (M. repens). Creep
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145Stachys 8lvensis. Field woundwor
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5cutellaria x hybrida (5. galericul
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Origanum vu/gare. Marjoram.Near Fis
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CALLlTRICHACEAE151Ca/litriche is a
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153Fraxinus excelsior. Ash.4 5Local
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~'Cymbalaria mural is. Ivy-leaved t
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157Veronica scutellata. Marsh speed
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159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73