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Flora of Ashdown Forest - Botanical Society of the British Isles

Flora of Ashdown Forest - Botanical Society of the British Isles

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fences or topography. It may dominate <strong>the</strong> woodland floor vegetation in <strong>the</strong> older woodlands, and <strong>the</strong> isolated clumpin Broadstone Warren is presumably a recent colonist.Very locally frequent in <strong>the</strong> Weald, especially in rocky woodlands and on acidic soils, but surely not on heaths assuggested by Hall (1980), It is commonest in nor<strong>the</strong>rn and western Britain and grows at high altitude in <strong>the</strong> Scottishmountains, usually on ungrazed ledges but also sometimes on open, grazed hillsides where <strong>the</strong> patches increasevegetatively but rarely flower. It is widespread in sou<strong>the</strong>rn, central and western Europe occurring mainly in <strong>the</strong>mountains in <strong>the</strong> eastern part <strong>of</strong> its range, and occurs in Asia Minor.197Luzula campestris. Field wood-rush, EasterMgrass, Good Friday grass.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Widespread in virtually every square on <strong>the</strong> <strong>Forest</strong>. It forms patches in short,unimproved, neutral and acidic grassland, on verges, playing fields, golf course fairways,and especially in lawns. The patches slowly increase in size and eventually <strong>the</strong> centresmay die leaving a ring. It flowers from March onwards, hence <strong>the</strong> name EasterMgrass,and <strong>of</strong>ten occurs with Carex caryophyllea, but is very difficult to find by midsummer as<strong>the</strong> leaves die down until <strong>the</strong> autumn. It is very tolerant <strong>of</strong> mowing and grazing and is<strong>of</strong>ten frequent in mossy, heavily rabbit-grazed areas.Common and widespread in Sussex and Britain. Widespread in Eurasia, and in <strong>the</strong>mountains <strong>of</strong> tropical Africa.Luzula multiflara sensu lata. Many-flowered woodMrush.In Britain L. multitlora s.l. is usually considered to consist <strong>of</strong> L. multitlora and L. congesta. L. congesta is an octoploidwhich is <strong>of</strong>ten treated as an infraspecific taxon <strong>of</strong> L. multiflora in Britain but it appears to have been misunderstooddue to poor characters being given in <strong>the</strong> floras (Kirschner & Rich 1993). The two species can be separated asfollows:L. multif/ora fEhrh.) Lej. subsp. multiflora: Inflorescences usually pedunculate. An<strong>the</strong>rs as long as, or up to two timeslonger than <strong>the</strong> filaments. Seeds oblong-ovoid, 0.9-1.1 (-1.2) mm long, usually 0.8 mm wide.L. congesta (Thuill.) Lej.: Inflorescence congested, some <strong>of</strong> <strong>the</strong> clusters may be pedunculate. An<strong>the</strong>rs as long as <strong>the</strong>filaments. Seeds ovoid, 1.2-1.5 mm long, 0.9-1.0 mm wide.All records for <strong>the</strong> segregates have been determined by TR from seed measurements. Records for <strong>the</strong> aggregate areei<strong>the</strong>r <strong>of</strong> vegetative plants or from o<strong>the</strong>r recorders.L. multif/ora subsp. multiflora is much rarer on <strong>the</strong> <strong>Forest</strong> and tends to occur on bare mineral soils in <strong>the</strong> open asaround Gills Lap. It is widespread in Europe, and is itself a complex species in Europe represented by a range <strong>of</strong>morphologically and/or karyologically distinct populations at various ploidy levels. A hexaploid cytotype (subsp.mu/tit/ora) occurs in continental Europe and Britain in a range <strong>of</strong> habitats from <strong>the</strong> lowlands to <strong>the</strong> mountains, mostlyin meadows and mesophilous woodlands. In geographically or ecologically marginal regions, diploid and tetraploidpopulations can be found, and a new tetraploid has recently been found in Ireland (Kirschner & Rich 1993, 1996).L. congesta is common on rides, in wet heaths and bogs, open woodland, and damp grassland on <strong>Ashdown</strong><strong>Forest</strong>, and seems to be <strong>the</strong> common widespread plant <strong>of</strong> most <strong>of</strong> Sussex. It is an endemic <strong>of</strong> western Europe. It<strong>of</strong>ten occurs on peaty organic soils.L. mu/tit/ora s.l. is common in <strong>the</strong> Weald in damp acidic habitats and is widespread in Britain.Luzula mu/t1'f/ora 5.1.L. mu/tit/ora subsp. multif/oraMany-flowered wood-rushL. congestaClustered wood-rush.3 -++--'L~_--'-+-4 5

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