Santander, February 19th-22nd 2008 - Aranzadi
Santander, February 19th-22nd 2008 - Aranzadi
Santander, February 19th-22nd 2008 - Aranzadi
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Oysters ancient and modern: potential shape variation with habitat in flat oysters (Ostrea edulis L.), and its possible use in archaeology<br />
181<br />
shapes would probably require analysis by ANCO-<br />
VA on the log-transformed data (Campbell <strong>2008</strong>:<br />
115-116). Shell shapes in the modern samples forming<br />
a transect across the Solent were compared<br />
by ANOVAs on the shape ratios, and shell shapes<br />
in the two archaeological morphotypes from deposit<br />
2239 were compared by t-tests on the shape<br />
ratios (Sokal and Rolph 1995: 223-225). Since<br />
sample sizes were small and the distributions of<br />
the ratios may not have been normal, distributionfree<br />
tests were also employed: shell shape ratios in<br />
the modern transect samples were subjected to<br />
Kruskal-Wallis tests (Sokal and Rolph 1995: 423-<br />
427), and the two archaeological morphotypes<br />
were compared by Mann-Whitney U tests (Sokal<br />
and Rolph 1995: 427-430). A difference was said<br />
to be significant if its probability was less than 0.05.<br />
4. RESULTS<br />
4.1. Relationship between dimensions<br />
The lines of maximum shell height (Hmax) and<br />
length (Lmax) were not consistently related to features<br />
of the shell. The point on the umbo which<br />
defined the dorsal point of maximum height could<br />
be above the chondrophore or either bourrelet,<br />
depending on the extent of curvature in the growth<br />
of the hinge. The point on the ventral edge defining<br />
Hmax could be anterior, directly ventrally or posterior<br />
to the adductor scar. However, the maximum<br />
dimensions in the plane of commissure did have<br />
consistent relationships with other shell features in<br />
both modern and archaeological shells, regardless<br />
of size or shape (Fig. 4). The line of maximum<br />
closure height (Hc) passed through the centre<br />
of the adductor muscle scar, usually within<br />
±1mm of the ventral edge of the scar; closure shell<br />
height and maximum body height are on the same<br />
alignment. Also, closure length Lc was parallel to<br />
the hinge axis, regardless of the relative orientation<br />
of Lc and Hc. These relationships held true even in<br />
one- or two-year-old shells, despite their xenomorphism<br />
(the strong trend for young shells to fit the<br />
shape of their substrates) (Stenzel 1971: 1021).<br />
The only shells which did not conform were those<br />
which bore clear evidence of trauma or physical<br />
growth restriction.<br />
4.2. Modern Oysters<br />
In the modern samples, the scatter in the relationship<br />
of height with other variables such as<br />
length (Fig. 5) was consistently broader with maximum<br />
shell height Hmax (Fig. 5a) than with closure<br />
height Hc (Fig. 5b). Regression coefficients (r 2 )<br />
were consistently lower with Hmax than with Hc for<br />
all shape ratios in all modern samples. For example,<br />
the relationship between height and length in<br />
the sample with the largest size range (Southsea<br />
Beach) had a greater scatter and larger regression<br />
coefficient with Hmax (Fig. 5a) than with Hc (Fig.<br />
5b). Distributions of shape ratios were also consistently<br />
broader in range and more poly-modal when<br />
generated using Hmax than when Hc was employed.<br />
For example, the distribution of shell HLR for<br />
Southsea Beach (Fig. 5c) was probably positively<br />
skewed and possibly poly-modal, while closure<br />
HLR (Fig. 5d) in the same sample was narrower in<br />
Figure 4. Inside of L. valve of O. edulis, showing measurements taken.<br />
Figure 5. Modern oyster O. edulis sample from Southsea Beach: height as a<br />
function of shell length for (a): shell height Hmax; (b): closure height Hc; and<br />
distribution of height-length ratio HLR using (c): shell height Hmax; (d): closure<br />
height Hc. Regression coefficient ‘ r2 ‘ for least-squares fitting to straight line.<br />
MUNIBE Suplemento - Gehigarria 31, 2010<br />
S.C. <strong>Aranzadi</strong>. Z.E. Donostia/San Sebastián