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Santander, February 19th-22nd 2008 - Aranzadi

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Oysters ancient and modern: potential shape variation with habitat in flat oysters (Ostrea edulis L.), and its possible use in archaeology<br />

181<br />

shapes would probably require analysis by ANCO-<br />

VA on the log-transformed data (Campbell <strong>2008</strong>:<br />

115-116). Shell shapes in the modern samples forming<br />

a transect across the Solent were compared<br />

by ANOVAs on the shape ratios, and shell shapes<br />

in the two archaeological morphotypes from deposit<br />

2239 were compared by t-tests on the shape<br />

ratios (Sokal and Rolph 1995: 223-225). Since<br />

sample sizes were small and the distributions of<br />

the ratios may not have been normal, distributionfree<br />

tests were also employed: shell shape ratios in<br />

the modern transect samples were subjected to<br />

Kruskal-Wallis tests (Sokal and Rolph 1995: 423-<br />

427), and the two archaeological morphotypes<br />

were compared by Mann-Whitney U tests (Sokal<br />

and Rolph 1995: 427-430). A difference was said<br />

to be significant if its probability was less than 0.05.<br />

4. RESULTS<br />

4.1. Relationship between dimensions<br />

The lines of maximum shell height (Hmax) and<br />

length (Lmax) were not consistently related to features<br />

of the shell. The point on the umbo which<br />

defined the dorsal point of maximum height could<br />

be above the chondrophore or either bourrelet,<br />

depending on the extent of curvature in the growth<br />

of the hinge. The point on the ventral edge defining<br />

Hmax could be anterior, directly ventrally or posterior<br />

to the adductor scar. However, the maximum<br />

dimensions in the plane of commissure did have<br />

consistent relationships with other shell features in<br />

both modern and archaeological shells, regardless<br />

of size or shape (Fig. 4). The line of maximum<br />

closure height (Hc) passed through the centre<br />

of the adductor muscle scar, usually within<br />

±1mm of the ventral edge of the scar; closure shell<br />

height and maximum body height are on the same<br />

alignment. Also, closure length Lc was parallel to<br />

the hinge axis, regardless of the relative orientation<br />

of Lc and Hc. These relationships held true even in<br />

one- or two-year-old shells, despite their xenomorphism<br />

(the strong trend for young shells to fit the<br />

shape of their substrates) (Stenzel 1971: 1021).<br />

The only shells which did not conform were those<br />

which bore clear evidence of trauma or physical<br />

growth restriction.<br />

4.2. Modern Oysters<br />

In the modern samples, the scatter in the relationship<br />

of height with other variables such as<br />

length (Fig. 5) was consistently broader with maximum<br />

shell height Hmax (Fig. 5a) than with closure<br />

height Hc (Fig. 5b). Regression coefficients (r 2 )<br />

were consistently lower with Hmax than with Hc for<br />

all shape ratios in all modern samples. For example,<br />

the relationship between height and length in<br />

the sample with the largest size range (Southsea<br />

Beach) had a greater scatter and larger regression<br />

coefficient with Hmax (Fig. 5a) than with Hc (Fig.<br />

5b). Distributions of shape ratios were also consistently<br />

broader in range and more poly-modal when<br />

generated using Hmax than when Hc was employed.<br />

For example, the distribution of shell HLR for<br />

Southsea Beach (Fig. 5c) was probably positively<br />

skewed and possibly poly-modal, while closure<br />

HLR (Fig. 5d) in the same sample was narrower in<br />

Figure 4. Inside of L. valve of O. edulis, showing measurements taken.<br />

Figure 5. Modern oyster O. edulis sample from Southsea Beach: height as a<br />

function of shell length for (a): shell height Hmax; (b): closure height Hc; and<br />

distribution of height-length ratio HLR using (c): shell height Hmax; (d): closure<br />

height Hc. Regression coefficient ‘ r2 ‘ for least-squares fitting to straight line.<br />

MUNIBE Suplemento - Gehigarria 31, 2010<br />

S.C. <strong>Aranzadi</strong>. Z.E. Donostia/San Sebastián

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