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5.6. Pelagic biota<br />

Within the existing contractor exploration claims, the environment<br />

includes expanses <strong>of</strong> oligotrophic subtropical ocean. Data were not<br />

available for the Indian Ocean claim area in particular, but the environment<br />

is similar <strong>and</strong> can be expected to have the same <strong>and</strong> similar types <strong>of</strong><br />

biological resources as those found in the CCFZ.<br />

5.6.1. Bacteria<br />

Bacteria are found throughout the water column as well as in the<br />

sediments. These microbial decomposers are concentrated at the sea<br />

surface, in the oxygen-minimum zone <strong>and</strong> in the sediments. They are<br />

associated, as organic aggregates, with all detritus, <strong>and</strong> their<br />

remineralisation <strong>of</strong> organic matter provides a major source <strong>of</strong> nutrients for<br />

use by algae. Sorokin 60 found that bacterial aggregates constituted 20-40%<br />

<strong>of</strong> oceanic particles, with little depth variability other than a slight<br />

concentration at the bottom <strong>of</strong> the thermocline. The size <strong>of</strong> the aggregates<br />

(larger than some small phytoplankton) makes them easily available as<br />

substrate for fine filter feeders, but also for coarse filter feeders <strong>and</strong> even<br />

selective feeders such as larval fish. He estimates that "bacterioplankton"<br />

are as important as algae as a primary food substrate.<br />

5.6.2. Phytoplankton<br />

El Sayed <strong>and</strong> Taguchi 61 examined water samples from the DOMES<br />

area <strong>and</strong> identified 163 types <strong>of</strong> diatoms, 122 din<strong>of</strong>lagellates, 48<br />

coccolithophorids <strong>and</strong> 15 other types. The coccolithophorid Gephyrocapsa<br />

huxleyi was the dominant species <strong>of</strong> phytoplankton at the depth <strong>of</strong> the<br />

chlorophyll maximum. In subtropical surface waters, nutrient<br />

concentrations are low <strong>and</strong> phytoplankton populations, measured by either<br />

cell counts or chlorophyll a concentrations, are sparse. Daily primary<br />

productivity is low, 100-200 mg <strong>of</strong> carbon per m 2 .<br />

A subsurface maximum is seen in chlorophyll a concentrations<br />

below the mixed layer. Concentrations <strong>of</strong> about 0.06 mg/m 3 are found in<br />

the mixed layer but are three to four times those at 70-80 m. Typically,<br />

production is limited by the low availability <strong>of</strong> nutrients in near-surface<br />

waters <strong>and</strong> by the decreasing availability <strong>of</strong> light at greater depths. Surface<br />

light at these latitudes is generally so intense that a thin layer at the surface<br />

experiences photoinhibition. The highest values <strong>of</strong> gross primary production<br />

INTERNATIONAL SEABED AUTHORITY 132

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