December 2012 Number 1 - Utah Native Plant Society

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Utah Native Plant Society Dam Mountains, mislabeled as Bear Valley and Beaver Valley by Gray], 20-28 Apr 1877 (fl, fr), Palmer s.n. (GH, K). ARIZONA: MOHAVE CO.: Virgin Narrows about 5 mi NE of Littlefield, 2 Jun 1977 (fl, fr), Gierisch 3954 (BRY, NY); Hedricks Canyon [Virgin Mountains], T40N R15W S23, 2 Apr 1981 (fl, imm fr), Gierisch & Morgart 4824 (BRY, NY). UTAH: WASH- INGTON CO.: Beaver Dam Mountains, Bull Dog Canyon, 25 Apr 2003 (fr), Alexander 1388 (OSC, UNLV), 6 May 2005 (fl,fr), Alexander 2120 (OSC,UNLV), 6 May 2005 (fl, fr), Alexander 2121 (OSC,UNLV), 6 May 2005 (fl, fr), Alexander 2127 (OSC,UNLV); Bulldog Knolls, 6 May 2006 (fl, fr), Alexander 2132 (OSC,UNLV), 6 May 2006, Alexander 2134 [individuals A,B,C] (OSC, UNLV), 6 May 2006 (fl, fr), Alexander 2135 (OSC, UNLV); Bull Dog Knolls, S slope of S knoll, T43S R18W S28, 1097 m, 21 Apr 1986 (fl, fr), Baird 2324 (BRY); Bulldog Knolls, north knoll, T43S R18W S21, 341 m, 28 Mar 1986 (fl, imm fr), Higgins & Barnum 16267 (BRY, NY); Bulldog Canyon, Beaver Dam Mountains, W slope, T43S R18W S26, 381 m, 15 Apr 1983 (fl, fr), Neese & Welsh 13062 (BRY, NY); Cedar Pockets wash, along a sequential drainage, T43S R17W S20, 1219 m, 5 May 1986 (fr), Welsh & Atwood 23742 (BRY). ACKNOWLEDGMENTS Numerous individuals provided advice, funding and encouragement during various stages of this project including: Wesley E. Niles; Aaron Liston, my thesis advisor; Richard Halse of the OSC Herbarium; Kathryn Birgy of the UNLV Herbarium; Arnold Tiehm and the Nevada Native Plant Society (and the NNPS Small Grants Program); Lisa Decesare of the Library of the Gray Herbarium, Harvard University; the Moldenke Fund for Plant Systematics and the Hardman Fund at Oregon State University; and my committee members, Mary Santelmann, David Lytle, Rich Cronn, and Paul Farber. Additional thanks go to the late R.C. Barneby and the Barneby Fund for Research in Legume Systematics, which funded a month of research at NY. The following herbaria's online databases were consulted for this project: CAS, DS, GH, JEPS, K, MO, NY, OSC, ORE, RSA, UC, UNLV, US, WILLU. I thank the curators and staff of the following herbaria for access to their collections or loans of their specimens: BRY, CAS, DS, GH, JEPS, K, NY, OSC, ORE, RM, RSA, UC, UNLV, US, WILLU. LITERATURE CITED Alexander, J.A. 2005. The taxonomic status of Astragalus mokiacensis (Fabaceae), a species endemic to the Southwestern United States. Brittonia 57:320-333. Alexander, J.A. 2008. A taxonomic revision of Astragalus mokiacensis and allied taxa within the Astragalus lentiginosus complex of Section Diphysi. Ph.D. dissertation. Oregon State University, Corvallis, OR. Alexander, J.A. 2009. The types of Astragalus Section Diphysi (Fabaceae), a complex endemic to Western North America, Part I: lectotypifications, epitypifications, and new combinations of several taxa. Journal of the Botanical Research Institute of Texas 3:211-218. Barneby, R.C. 1945. Pugillus Astragalorum - IV: the section Diplocystium. Leaflets of Western Botany 4: 65- 147. Barneby, R.C. 1956. Pugillus Astragalorum - XVII: four new species and one variety. Leaflets of Western Botany 8: 14-23. Barneby, R.C. 1964. Atlas of North American Astragalus. (2 volumes). Memoirs of the New York Botanical Garden. 13:1-1188. Barneby, R.C. 1989. Intermountain Flora. Volume III Part B. Eds. A. Cronquist, A.H. Holmgren, N.H. Holmgren, J.L. Reveal, P.K. Holmgren. The New York Botanical Garden. Bronx, New York. 279 pp. Cracraft, J. 1983. Species concepts and speciation analysis. Current Ornithology 1: 159-187. Easdale, T.A., Gurvich, D.E., Sersic, A.N. & Healey, J.R. 2007. Tree morphology in seasonally dry montane forest in Argentina: Relationships with shade tolerance and nutrient shortage. Journal of Vegetation Science 18: 313-326. Gower, J.C. 1971. A General Coefficient of Similarity and Some of Its Properties. Biometrics 27:857-871. Gray, A. 1849. Plantae Fendlerianae Novi-Mexicanae. Memoirs of the American Academy of Arts and Sciences. Series 2. 1:1-116. Gray, A. 1863. A Revision and Arrangement (mainly by the fruit) of the North American species of Astragalus and Oxytropis. Proceedings of the American Academy of Arts and Sciences. 6:188-237. Gray, A. 1865. Characters of some new plants of California and Nevada, chiefly from the collections of Professor William H. Brewer, botanist of the State Geological Survey of California, and of Dr. Charles L. Anderson, with revisions of certain genera or groups. Proceedings of the American Academy of Arts and Sciences. 6:519-556. Gray, A. 1878. Contributions to the botany of North America. Proceedings of the American Academy of Arts and Sciences. 13:361-374. Hammer, O., Harper, D.A.T., and P.D. Ryan. 2001. PAST: Palaeontological Statistics software package for education and data analysis. Palaeontological Electronica 4:9 pp. Hooker, W.J. 1831. Flora Borelli-American or the botany of the northern parts of British America. 2 (12):97-160. 150
Calochortiana December 2012 Number 1 Isely, D. 1998. Native and naturalized Leguminosae (Fabaceae) of the United States. Brigham Young University. Provo, Utah. 1007 pp. Jones, M.E. 1898. Contributions to Western Botany 8. Contributions to Western Botany. 8:1-43. Jones, M.E. 1923. Revision of North American species of Astragalus. Text distributed Feb. 15, 1923; Plates, June 20, 1923. Salt Lake City, Utah. 288 pp. Kearney, T.H. and R.H. Peebles. 1960. Arizona Flora. University of California Press. Berkeley, California. 1079 pp. Knaus, B.J. 2008. A Fistful of Astragalus: phenotypic and genotypic basis of the most taxon rich species in the North American Flora. Ph.D. dissertation. Oregon State University, Corvallis, OR. Luckow, M. 1995. Species Concepts: assumptions, methods, and applications. Systematic Botany. 20:589- 605. Munz, P.A. and D.D. Keck. 1959. A California flora. University of California Press. Berkeley, California. Nixon, K.C., and Q.D. Wheeler. 1990. An amplification of the phylogenetic species concept. Cladistics 6: 211-223. Rydberg, P.A. 1929. Astragalinae. North American Flora 24(5): 251-462. Sokal, R. R. 1973. The species problem reconsidered. Systematic Zoology. 22: 360-374. Swofford, D. L. 2002. PAUP*: phylogenetic analysis using parsimony. ver. 4 beta 10. Sinauer Associates, Inc., Sunderland, Massachusetts. Watson, S. 1871. Botany. In C. King, Report of the geological exploration of the fortieth parallel, Vol. 5. Washington: Government Printing Office. 525 pp. Welsh, S.L. 1978. Utah flora: Fabaceae (Leguminosae). Great Basin Naturalist. 38:225-367 Welsh, S.L. 1993. Leguminosae (Fabaceae). Pp 379- 463 in S. L . Welsh, N. D. Atwood, S. Goodrich, and L. C. Higgins, eds., A Utah Flora. 2nd ed. Monte L. Bean Life Science Museum, Provo, Utah. Welsh, S.L. 2003. Leguminosae (Fabaceae). Pp 350- 427 in S. L . Welsh, N. D. Atwood, S. Goodrich, and L. C. Higgins, eds., A Utah Flora. 3rd ed. Monte L. Bean Life Science Museum, Provo, Utah. Welsh, S.L. 2007. North American species of Astragalus (Leguminosae): A taxonomic review. Monte L. Bean Life Science Museum. Provo, Utah. 932 pp. Welsh, S.L., and N.D. Atwood. 2001. New taxa and nomenclatural proposals in miscellaneous families- Utah and Arizona. Rhodora. 103: 71-95. Wojciechowski, M. F., M. J. Sanderson, B. G. Baldwin, & M. J. Donoghue. 1993. Monophyly of aneuploid Astragalus (Fabaceae): evidence from nuclear ribosomal DNA internal transcribed spacer sequences. American Journal of Botany. 80: 711-722. Wojciechowski, M.F., M.J. Sanderson, J.-M. Hu. 1999. Evidence on the monophyly of Astragalus (Fabaceae), and its major subgroups based on nuclear ribosomal DNA ITS and chloroplast DNA trnL intron data. Systematic Botany. 24: 409-437. APPENDIX 1 List of vouchers used in morphometric analyses not cited in the Taxonomic Revision Note: All vouchers listed herein have been seen by the author. The "!" designation is not used herein as a result. Astragalus lentiginosus var. ambiguus Barneby U.S.A. ARIZONA: MOHAVE CO.: Peach Springs, 11 May 1941, Ripley & Barneby 3403 (RSA). Astragalus lentiginosus var. araneosus (Sheld.) Barneby U.S.A. UTAH: MILLARD CO.: Frisco, June 1880, Jones 1807 [June 1880, Jones s.n., interpreted herein as an isotype] (NY, ORE, GH). Astragalus lentiginosus var. bryantii (Barneby) J.A. Alexander U.S.A. ARIZONA: COCONINO CO.: head of Phantom Canyon in Grand Canyon, 15 December 1939, Bryant s.n. (CAS); 10 yds N of Colorado River, 11 mi S of Phantom Ranch, directly N of Grand Canyon Village, 11 April 1960, Crosswhite 642 (NY); at mouth of Hermit Creek, in sand, Grand Canyon of the Colorado River, 10 April 1917, Eastwood 5991 (GH); "Utah Flat", Grand Canyon N.P., ca. 0.83 mi NW of Phantom Ranch and Bright Angel Creek, 09 April 1993, Hodgson & Anderson 2085 (NY); Colorado River, Bass Rapids, 108 miles below Lees Ferry, 01 May 1971, Holmgren, et al. 15502 (NY); Colorado River, Grand Canyon near confluence of Clear Creek, 3.5 miles upriver from Kaibab Suspension Bridge (near Phantom Ranch), 1 mile up Clear Creek Canyon, 08 May 1971, Holmgren, et al. 15609 (GH, NY); Colorado River, Grand Canyon, Shinumo Creek, 108.5 river mi below Lees Ferry, 17 air mi NW of Grand Canyon Village, 10 May 1971, Holmgren, et al. 15615 (NY); Grand Canyon N.P., at confluence of Bright Angel Creek and Colorado River, 20 March 1968, Spellenberg 1826. Astragalus lentiginosus var. iodanthus (S. Watson) J.A. Alexander U.S.A. NEVADA: CARSON CITY (ORMSBY) CO.: Empire City [Carson City vicinity], 19 May 1882, Jones 3837 [May 1882, Jones s.n., interpreted herein as a duplicate] (NY, ORE). 151
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