Review

∫∑§«“¡
«“√ “√«‘∑¬“»“ µ√å ¡¢. ªï∑’Ë 39 ©∫—∫∑’Ë 1 39
96
100
100
0.01
100
Halococcus saccharolyticus JCM 8878 T (AB004876)
Halococcus dombrowskii JCM 12289 T (AJ420376)
100
Halococcus dombrowskii JCM 12289 T (AJ420376)
Halococcus qingdaogense JCM 13587 T (AY243109)
54 Halococcus thailnadensis
Natrinema gari (AB289741)
Natrinema pellirubrum JCM 10476 T (AJ002947)
96 Natrinema pallidum JCM 8980 T (AJ002949)
Halobacterium piscisalci (AB285020)
62
Halobacterium salinarum DSM 3754 T (AJ496185)
100
Halobacterium salinarum strain DS2-5
52 Haloarcula amylolytica JCM 13557 T rrnB (DQ826513)
95
Haloarcula salaria rrnA (FJ429317)
Haloarcular tradensis rrnB (FJ429316)
78
60 Haloarcula amylolytica JCM 13557 T rrnC (DQ854818)
Haloarcula quadrata 801030/1 T rrnB (EF645694)
64
70
Haloarcula salaria rrnB (FJ429318)
Haloarcula tradensis rrnC (FJ429314)
Haloarcula tradensis rrnA (FJ429313)
Haloarcula quadrata JCM 11048 T rrnA (AB010965)
99 Haloarcula amylolytica JCM 13557 T rrnA (DQ826512)
Natronobacterium pharaonis JCM 8858 T (D87971)
√Ÿª∑’Ë 2 · ¥ß·ºπ¿Ÿ¡‘µâπ‰¡â¢ÕßÕ“√凧’¬Õ∫‡§Á¡ “¬æ—π∏ÿå„À¡à∑’˧—¥·¬°®“°ª√–‡∑»‰∑¬ (∑’Ë¡“: Namwong et al.,
2007; Namwong et al., inpress; Yachai et al., 2008; Tapingkae et al., 2008)
°“√ª√—∫µ—«¢ÕßÕ“√凧’¬Õ∫‡§Á¡„π ¿“«–∑’Ë¡’
‡°≈◊Õ (Osmotic adaptation of halophilic
archaea)
„π ¿“«– ¡¥ÿ≈ (osmotic balance) √–À«à“ß
§«“¡‡¢â¡¢âπ¢Õß “√≈–≈“¬¿“¬„π (intracellular
solutes, S) °—∫¿“¬πÕ°‡´≈≈å∑”„À⇴≈≈å®ÿ≈‘π∑√’¬å
“¡“√∂‡®√‘≠‡µ‘∫‚µ‰¥â·≈–§ß√Ÿª√à“߇¥‘¡ (√Ÿª∑’Ë 3.1)
„π ¿“«–∑’Ë¡’‡°≈◊Õ Ÿß (hypertonic condition) §«“¡
‡¢â¡¢âπ¢Õß “√≈–≈“¬¿“¬πÕ°‡´≈≈å¡’¡“°°«à“¿“¬„π
‡´≈≈å®ÿ≈‘π∑√’¬å ∑”„ÀâπÈ”¿“¬„π‡´≈≈å‰À≈ÕÕ°πÕ°‡´≈≈å
‡æ◊ËÕª√—∫ ¡¥ÿ≈√–À«à“߇´≈≈å°—∫ ‘Ëß·«¥≈âÕ¡ àߺ≈„Àâ
‡´≈≈å Ÿ≠‡ ’¬πÈ” (dehydration) ®π„π∑’Ë ÿ¥‡´≈≈剡à
“¡“√∂‡®√‘≠‡µ‘∫‚µ (√Ÿª∑’Ë 3.2) ”À√—∫Õ“√凧’¬Õ∫
‡§Á¡ “¡“√∂‡®√‘≠‰¥â¥’„π ¿“«–¥—ß°≈à“«‡π◊ËÕß®“°¡’
°√–∫«π°“√ª√—∫µ—«„π ¿“«–·«¥≈âÕ¡∑’Ë¡’‡°≈◊Õ ¥—ß
µàÕ‰ªπ’È
1. ‡æ‘Ë¡ª√‘¡“≥‰ÕÕÕπ “√Õπ‘π∑√’¬å
(Accmulation of inorganic ions)
‡àπ Halobacterium salinarum µ√«®æ∫
‚´‡¥’¬¡‰ÕÕÕπ (Na + ) ¿“¬πÕ°‡´≈≈åª√–¡“≥ 4.0
‚¡≈“√å ·µà¡’ª√‘¡“≥‚æ·∑ ‡´’¬¡‰ÕÕÕπ¿“¬„π‡´≈≈å
¡“°∂÷ß 4.57 ‚¡≈“√å (Martin et al., 1999; Christian