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Impact of Climate Change on Arab Countries - (IPCC) - Working ...

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ARAB ENVIRONMENT: CLIMATE CHANGE 107<br />

and mountains in the <strong>Arab</strong> world is c<strong>on</strong>strained<br />

by water availability, temperature, and/or their<br />

substrate (Table 1). The potential <str<strong>on</strong>g>of</str<strong>on</strong>g> these<br />

ecosystems to resp<strong>on</strong>d to climate change by<br />

migrati<strong>on</strong> is therefore limited and their survival<br />

more doubtful.<br />

IV. ECOSYSTEM COMPOSITION<br />

AND SPECIES VULNERABILITY<br />

TO CLIMATE CHANGE<br />

Adaptati<strong>on</strong>s to climate change will alter entire<br />

ecosystems in terms <str<strong>on</strong>g>of</str<strong>on</strong>g> physical, chemical and<br />

biological features and/or alter species compositi<strong>on</strong><br />

forcing species to disperse, adapt or face<br />

ultimate extincti<strong>on</strong>. Wherever c<strong>on</strong>diti<strong>on</strong>s favour<br />

large changes in the number <str<strong>on</strong>g>of</str<strong>on</strong>g> species that coexist<br />

in a given area, or species richness, communities<br />

can be expected to undergo major reorganizati<strong>on</strong>.<br />

As a result protecti<strong>on</strong> <str<strong>on</strong>g>of</str<strong>on</strong>g> representative<br />

and/or ecosystems currently in existence<br />

will be problematic (Currie, 2001). Since<br />

species are generally expected to resp<strong>on</strong>d individually<br />

to climate change, there is potential for<br />

novel species combinati<strong>on</strong>s to occur, and for<br />

present day relati<strong>on</strong>ships to become increasingly<br />

decoupled (Thuiller et al., 2006). For instance<br />

the loss <str<strong>on</strong>g>of</str<strong>on</strong>g> the rarest species can be compensated<br />

by increased growth <str<strong>on</strong>g>of</str<strong>on</strong>g> the dominant species,<br />

whereas reducti<strong>on</strong>s in the density <str<strong>on</strong>g>of</str<strong>on</strong>g> the dominant<br />

species cannot be compensated by rare<br />

species.<br />

In terms <str<strong>on</strong>g>of</str<strong>on</strong>g> species richness, although many studies<br />

show positive effects <str<strong>on</strong>g>of</str<strong>on</strong>g> biodiversity <strong>on</strong> ecosystem<br />

functi<strong>on</strong>, others do not. The reas<strong>on</strong> may be<br />

related to the positi<strong>on</strong> <str<strong>on</strong>g>of</str<strong>on</strong>g> the species most affected<br />

by climate change. For instance, if the loss <str<strong>on</strong>g>of</str<strong>on</strong>g><br />

species is at top trophic levels or a keyst<strong>on</strong>e<br />

species, then it may have particularly str<strong>on</strong>g<br />

ecosystem effects. On the other hand climate<br />

change induced loss <str<strong>on</strong>g>of</str<strong>on</strong>g> species at different trophic<br />

levels or different species groups will be difficult<br />

to predict in resp<strong>on</strong>se to climate change. It is<br />

quite possible, however, that over the short term<br />

(decades to centuries) species richness will<br />

decrease, even in areas where richness is predicted<br />

to increase in the l<strong>on</strong>g term: as climate<br />

changes, species that are intolerant <str<strong>on</strong>g>of</str<strong>on</strong>g> local c<strong>on</strong>diti<strong>on</strong>s<br />

may disappear relatively quickly while<br />

migrati<strong>on</strong> <str<strong>on</strong>g>of</str<strong>on</strong>g> new species into the area may be<br />

quite slow (Currie, 2001).<br />

Understanding species dispersal habits becomes<br />

essential since, with respect to the timescale c<strong>on</strong>sidered<br />

by climate change predicti<strong>on</strong>s, dispersal<br />

habits can range from those that would be unable<br />

to disperse significantly to those that could readily<br />

disperse and establish. For example, species<br />

with large seeds will not migrate as quickly as<br />

species with lighter seeds or those seeds that can<br />

be dispersed by animals. Some simulati<strong>on</strong> studies,<br />

however, show that many <str<strong>on</strong>g>of</str<strong>on</strong>g> the species<br />

whose potential distributi<strong>on</strong> ranges change significantly<br />

will become n<strong>on</strong>-viable because <str<strong>on</strong>g>of</str<strong>on</strong>g><br />

delays in populati<strong>on</strong> resp<strong>on</strong>ses to climate change<br />

(Miles et al., 2004). In general, species that currently<br />

have a broad distributi<strong>on</strong> range will most<br />

likely have high dispersal ability while those that<br />

exist in few sites will likely have low dispersal<br />

ability and be more threatened by climate change<br />

driven disasters such as fires, droughts, and pest<br />

outbreaks. On the other hand, migrating species<br />

could experience multiple extincti<strong>on</strong>s as they<br />

encounter anthropogenic land transformati<strong>on</strong>s<br />

or simply reach the mountain summit or water<br />

body edges.<br />

Future predicti<strong>on</strong>s <str<strong>on</strong>g>of</str<strong>on</strong>g> species vulnerability will<br />

need to be based <strong>on</strong> analytical tools such as broad<br />

modelling frameworks that focus <strong>on</strong> the species<br />

suitable climate space through the use <str<strong>on</strong>g>of</str<strong>on</strong>g> bioclimatic<br />

envelop models which depend <strong>on</strong> the<br />

analysis <str<strong>on</strong>g>of</str<strong>on</strong>g> the complex interacti<strong>on</strong>s at different<br />

levels (Pears<strong>on</strong> and Daws<strong>on</strong>, 2003) and/or integrative<br />

sensitivity analysis modelling which<br />

examines the effect <str<strong>on</strong>g>of</str<strong>on</strong>g> climate change <strong>on</strong> individual<br />

species, especially those that are rare, threatened<br />

and climate sensitive as well as ecological<br />

processes (Hannah et al., 2002).

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