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90<br />

shape. Colony obviously dead when collected, the coenosteum<br />

well worn and discolored to a light brown. Branches robust,<br />

basal branches about 31 mm in diameter, distal blunt tipped<br />

branches about 13 mm in diameter, branch anastomosis<br />

common. Corallites 5-6 mm in diameter and flush with<br />

coenosteum, perhaps due to wear of corallum. Theca vermiculatc<br />

as well as obscurely costate, the coenosteum penetrated by<br />

numerous small (0.32 mm in diameter) circular pores (?borings).<br />

Septa hexamerally arranged in 4 cycles in a Pourtales<br />

Plan, some of the larger corallites having several pairs of S5. S,<br />

slightly wider than S2, both cycles of septa attaining the<br />

columella. S3 narrowest of septa, each S3 flanked by a pair of<br />

S4 which fuse before the S3 and extend to the columella. Fossa<br />

shallow, containing a large, spongy columella.<br />

DISCUSSION.—Dendrophyllia cribrosa is treated as a member<br />

of the first group of species as defined in the generic<br />

discussion because it has an arborescent corallum and<br />

nonsympodially budded corallites. It is relatively easily<br />

distinguished from other North Pacific species by having thick,<br />

anastomotic branches; very short (flush) corallites oriented<br />

perpendicular to parent branches; circular corallites 4.5-8.0<br />

mm in diameter, and a robust columella. The proper authorship<br />

of the species is Milne Edwards and Haime (1851), not Milne<br />

Edwards and Haime (1860), as suggested by some authors.<br />

Blainville's earlier names published in lists must be considered<br />

as nomina nuda and de Haan's Oculina anastomozans was<br />

apparently only another unpublished manuscript name cited by<br />

Blainvilic (1830).<br />

MATERIAL EXAMINED.—New Records: None. Previous<br />

Records: Larger syntype of D. cribrosa. RMNH.<br />

TYPES.—Two syntypes (Plate 38tf./i) of D. cribrosa are<br />

deposited at the RMNH: the larger specimen described above<br />

(RMNH 9212. manuscript type of Dentipora anastomozans)<br />

and a much smaller fragment 8.2 cm long (RMNH 9209,<br />

manuscript type of Dentipora cribrosa). Type Locality:<br />

Unknown.<br />

DISTRIBUTION.—Japanese region: Sagami Bay. Honshu;<br />

?Sea of Japan off eastern and southern South Korea (Song,<br />

1982); Pleistocene of Kanagawa-ken; 7-40 m. Elsewhere:<br />

Off Banka. southeastern Sumatra; ?off Angola (Cabinda)<br />

and the Congo; 20-30 m.<br />

Dtndrophyllia japonic a Rehberg, 1892<br />

DendropMvllui faponuu Rehberg. IR92.28-29. pi. 4: fig. 4.—Van der Horst.<br />

1926 44-45. p| V figs. 4. V-Yabe and Eguchi. 1932a: 388; 1941b: 102 —<br />

Eguchi. 1934:367; I965295. 2 figs.. I968C6I-62. p». C13: figs. 3-5<br />

[synonymy].—Squires and Key**. 1967:28. pi 6: Tigs. 6-8 (synonymy).<br />

Not DrmJrvpMvllui japnmua Van der Honl. I922:.M. pi. 7: fig. 6 [junior<br />

homonym: replacement name D hnuhmai Van der Horst. 1926).<br />

DIAGNOSIS.—Colonies consist of a primary axial corallite up<br />

to 9 cm in height and up to 27 mm in GCD from which several<br />

slightly smaller, but still quite robust, corallites bud at irregular<br />

intervals. All colonies examined as well as those reported in the<br />

literature arc broken basally. but it would appear from the larger<br />

diameter of the basal fractures that the normal condition of the<br />

SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY<br />

species is attached, not free as in Eguchipsammia. Costae are<br />

flat and porous; epitheca absent. Septa hexamerally arranged in<br />

a Pourtales Plan, larger corallites having a full fifth cycle of<br />

septa (96), smaller corallites usually having numerous pairs of<br />

S5. Su2 equal in size, slightly exsert, and quite wide, their<br />

smooth inner edges bordering the columella. S3 and S4<br />

progressively smaller. Pairs of S5 fuse before their adjacent S4<br />

and again before the S3 in their half-system before reaching the<br />

columella. Fossa deep, containing an elongate narrow columella.<br />

DISCUSSION.—Dendrophyllia japonica belongs to the "second<br />

group" of Dendrophyllia species, those characterized as<br />

bushy, having small colonies with sparsely budded large<br />

corallites. It is compared to D. arbuscula in the account of that<br />

species.<br />

MATERIAL EXAMINED.—New Records: None. Previous<br />

Records: Specimens reported by Squires and Keyes (1967)<br />

from off New Zealand, NZOI.<br />

TYPES.—Two syntypes of Rehberg's D. japonica were<br />

present at the ZMB in the 1920's (Van der Horst, 1926), but it<br />

is likely that they were subsequently lost or destroyed. Type<br />

Locality: "Japan," depth unknown.<br />

DISTRIBUTION.—Japan: Sea of Japan off northwestern<br />

Honshu from Tsugaru Strait to Wakasa Bay; Sagami Bay;<br />

115-250 m. Elsewhere: Banda Sea; North Island, New<br />

Zealand; 114-549 m.<br />

Dendrophyllia arbuscula Van der Horst, 1922<br />

PLATE 38/-/<br />

? Dendrophyllia conferta — Rehberg, 1892:28.<br />

Dendrophyllia arbuscula Van der Horst. 1922:53. pi. 8: fig. 6.—Yabe and<br />

Eguchi. I932a:388; 1941b: 102.—Eguchi, 1934:367; 1965:294. 2 figs.;<br />

1968:C55-56. pi. C21: figs. 5, 13.—Not Kikuchi, 1968:9, pi. 5: fig. 6 [=<br />

Cladopsammia sp.].—Not Utinomi, 1971:220 [=Cladopsammia gracilis].—<br />

Eguchi and Miyawaki. 1975:54, pi. 6: fig. 4.—? Song, 1991:136, pi. 1: fig.<br />

4; pi. 3: fig. 1.<br />

^Dendrophyllia mbtornigera Eguchi. 1934:367 [nomen nudum]; 1968:C64,<br />

pi. C32: figs. 3. 4.<br />

^Dendrophyllia xufxornigera cylindrica Eguchi. 1934:367 [nomen nudum];<br />

l968:C64-65. pi. C32: figs. 1.2.<br />

^Dendrophyllia sp—Eguchi. I968:C67, pi. C30: figs. 6. 7.<br />

Tuhastraea cYK

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