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NUMBER 557 63
ony and 1 corallite, USNM 92705; TM (KT9202, YT2), 1
corallite, USNM 92706.
Types.—The holotype is deposited in the Biological Laboratory
of the Imperial Household, Tokyo (#657). Type Locality:
Amadaiba, Okino-Kannonzukadashi, Sagami Bay, 60-
80 m.
DISTRIBUTION.—Known only from off Japan: Sagami Bay;
Kagoshima Bay, Kyushu; Colnett Strait, Osumi Shoto,
northern Ryukyu Islands; 60-98 m.
Dasmosmilia Pourtales, 1880
DIAGNOSIS.—Solitary, ceratoid to turbinate, free. Parricidal
budding common. Theca and septa very thin. Paliform lobes
present before penultimate cycle of septa and occasionally also
before lower cycle septa. Columella trabecular. Endotheca
sparse or absent.
TYPE SPECIES.—Parasmilia lymani Pourtales, 1871, by
subsequent designation (Wells, 1933).
Dasmosmilia pacifica (Yabe and Eguchi, 1932), comb. nov.
PLATES 27/-I, 4 \f,g
Goniocyathus pacificus Yabe and Eguchi, 1932a:389, text-fig. 2; 1932b:443;
1942b: 122, 152-153, pi. 10: figs. 15, 16.—Eguchi, 1965:286, 2 figs.—
Zibrowius, 1980:70.
Caryophyllia pacifica—Wells, 1956:F422.
DESCRIPTION.—Corallum fragile, due to extremely thin
theca and septa. AH specimens examined had asexually
generated from a wedge-shaped fragment of parent corallum,
often resulting in somewhat irregular septal development and
symmetry. Largest specimen examined (syntype) 14.9 mm in
GCD. Corallum above base ceratoid, with a circular to
irregularly shaped calice. Each primary septum, along with its
2 adjacent higher-cycle septa, are fused at calicular edge and
project 1.0-1.5 mm above the edge, producing a serrate
margin. Costae slightly convex and coarsely granular, the
12-24 primary costae ridged. Corallum white, with a slight
brownish tint near the calice.
Septal symmetry irregular, the number of septa apparently
correlated with calicular diameter. Among the few specimens
examined, there are coralla with 12-24 primary septa, 12-24
secondary septa, 24-48 tertiary septa, one corallum (large
syntype, TIUS 50086) also having a pair of quaternary septa,
for a total of 98 septa. Primary septa exsert as much as 2.2 mm
and have vertical, sinuous inner edges, several of which clearly
bear tall twisted fascicular columella elements. Secondary
septa least exsert and only about 40% width of a primary. Each
secondary septum bears a wide paliform lobe (about 1.3 mm),
together forming a crown of 12-18 lobes. Tertiary septa
slightly more exsert but less wide than secondaries, becoming
rudimentary lower in fossa. When pairs of quaternary septa
occur within a half-system, they resemble tertiaries in size, and
the flanked tertiary is slightly enlarged in width and adds a
paliform lobe. Endothecal dissepiments not observed.
DISCUSSION.—Vaughan and Wells (1943) and Wells (1956)
placed the monotypic Goniocyathus Yabe and Eguchi, 1932a in
synonymy with Caryophyllia; however, Zibrowius (1980)
suggested that it might fall closer to Dasmosmilia. After
comparing the Japanese specimens reported herein with typical
D. lymani, I agree with Zibrowius that Goniocyathus is a junior
synonym of Dasmosmilia. In fact, the differences between D.
pacifica and D. lymani are quite minor. Dasmosmilia pacifica
differs in having exsert primary septa, always fused with
adjacent higher-cycle septa to form a serrate calicular margin,
whereas the primary septa of D. lymani are not very exsert and
not fused with adjacent septa near the calice. Also, the corallum
of D. pacifica is more open and the septa more widely spaced
than in D. lymani.
The thin theca of this genus facilitates its mode of asexual
budding.
MATERIAL EXAMINED.—New Records: Alb-3738, 1,
USNM 92710; Alb-5055, 1, USNM 92711; YO70-1009, 1,
ORI. Previous Records: 3 syntypes, TIUS.
TYPES.—The syntypes of G. pacificus are deposited at the
TIUS (50086, 50097)(Plate 4\f,g). Type Locality (vide Yabe
and Eguchi, 1942b): Soyo Maru-198: 34°17'45"N,
137 o 04'45 / ^(off Ise Bay, Honshu), 168 m.
DISTRIBUTION.—Known only from the Pacific coast of
Japan from off Boso Hanto, Chiba-ken, Honshu to southeastern
Kyushu; 168-355 m. Pleistocene of Ryukyu Islands (Yabe and
Eguchi, 1932b).
Goniocorella Yabe and Eguchi, 1932a
DIAGNOSIS.—Colonial, extratentacular budding forming
bushy colonies. Branch anastomosis common, the integrity of
the corallum further increased by numerous slender, tubular
coenosteal bridges. No pali or columella. Tabular endothecal
dissepiments common and widely spaced.
TYPE SPECIES.—Pourtalosmilia dumosa Alcock, 1902, by
original designation.
Goniocorella dumosa (Alcock, 1902)
PLATE 27/
Pourtalosmilia dumosa Alcock, 1902c:36-37, pi. 5: fig. 33.
Goniocorella dumosa.—Yabe and Eguchi, 1932a:389-390; 1936:167;
1943:494-496, figs. 1, 2; 1942b:162, 163.—Eguchi, 1965:291, 2 figs.;
1968:C43, pi. C9: figs. 11. 12.—Cairns, 1982:31-34, pi. 9: figs. 7-9; pi. 10:
figs. 1. 2 [synonymy].—Cairns and Keller. 1993:250. fig. 6E.—Song,
1991:134-135, pi. 1: fig. 3; pi. 2: figs. 4-7.
Goniocorella sp. aff. G. dumosa.—Eguchi and Miyawaki. 1975:58.
DESCRIPTION.—Colony bushy, budding often at right angle
to parent branch. Colony reinforced by slender (about 2 mm in
diameter), hollow, tubular coenosteal bridges, which unite
adjacent branches. Branches cylindrical, 3-5 mm in diameter,
each bearing a terminal calice of equal diameter. Costae
inconspicuous. Corallum white or light brown.