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NUMBER 557 91
(0.8-0.9 mm), with smooth, vertical inner edges that reach the
columella only low in fossa. S2 less exsert and slightly less
wide than an S,, but also have smooth inner edges that almost
attain the columella. S3 about half width of an S2. Each pair of
S4 within a half-system bend towards each other, resulting in
either a loose fusion of their inner edges before their enclosed
S3 or a closely parallel arrangement leading to the columella.
Inner edges of S4 slightly porous and laciniate to dentate, the
S^ being solid septa with entire inner edges. Fossa of
moderate to shallow depth, containing a well-developed,
discrete, elongate columella, sometimes constricted into 3
lobes by inner edges of the 4 lateral Sv Columella composed of
small, flattened, swirled elements that are tightly fused
together.
DISCUSSION.—Dendrophyllia arbuscula belongs to the
"second group" of Dendrophyllia species described in the
generic discussion: species having small, bushy colonies, with
relatively few large corallites. The only other species in this
group from the Japanese region is D. japonica Rehberg, 1892,
which differs in having larger corallites, more septa, and a less
developed columella.
Dendrophyllia subcornigera Eguchi, 1968 is only tentatively
synonymized with D. arbuscula, as the type and only
reported specimen of this species is not available for study. Its
description and illustrations are consistent with a large,
well-developed colony of D. arbuscula, much like that of the
holotype; in his original description, Eguchi (1968) noted the
resemblance. Likewise, D. subcornigera cylindrica Eguchi,
1968, which is also known from only one specimen, is also
tentatively synonymized but this specimen is even more robust
with long internodes between budded corallites and quite long
secondary corallites. Its growth form resembles that of the "first
group" of species of Dendrophyllia.
MATERIAL EXAMINED.—New Records: Alb-3707, 6,
USNM 22057, 2, ORI; Alb-3716, 4, USNM 22054; Alb-3718,
1. USNM 22053; Alb-3720, 2, USNM 22052; Alb-3730, I,
USNM 22056; Alb-3746, 2, USNM 22055; Alb-4935. 1,
USNM 92853; Alb-5068, 4, USNM 92854; Alb-5070, 2,
USNM 92855; TM (KT7811, OT6-2), 1, USNM 92856;
Okinosc, Sagami Bay, 183 m, Mortensen's Pacific Expedition,
15 June 1914, 3, ZMC.
TYPES.—Three syntypes of D. arbuscula are deposited at the
ZMA (Coel. 1254, 5477). Type Localities: Siboga-260. 277:
Banda Sea, 45-90 m.
The holotype of D. subcornigera is deposited at the
Biological Laboratory of the Imperial Household, Tokyo
(specimen number 896, reg. no. 40859). Type Locality:
Enoura Bay, Shizuoka ken. Honshu, depth unknown.
The holotype of D. subcornigera cylindrica is deposited at
the Biological Laboratory of the Imperial Household, Tokyo
(specimen number 897). Type Locality: Off Seto, Kanayamamaru,
Nishimuragun, Wakayama-ken; depth unknown.
DISTRIBUTION.—Japan: Sagami and Suruga Bays, Honshu;
Kii Strait; off Amakusa Islands; off Cheju Do, East China Sea;
40-240 m. Elsewhere: Banda Sea; 45-90 m.
Dendrophyllia boschmai Van der Horst, 1926
Dfndrophytlia japonica Van der Horst, 1922:51, pi. 7: fig. 6 [junior primary
homonym of I) japonica Rehberg. 1892].
Dendrophyllia htmhmai Van der Horst. 1926:44 [replacement name for D.
japonica Van der Horst. 1922].—Eguchi, 1934:367—Yahe and Eguchi.
1936:167; I968:C56-S7, pi. C15: figs. 4. 7; pi. CI6: figs. 3. 4; pi. C17: figs.
12, 15; pi. C30: fig. 1—Song, 1982:138, pi. 4: figs. 3, 4; 1991:136.
Dendrophyllia cyathoheloides Eguchi, 1934:367 [nomen nudum)
Dtndrophyllia boschmai cyathoneloides Eguchi. 1965:294, I fig.
Dfndrophytlia boschmai cyathohelioides [sic].—Eguchi. 1968:C57. pi. C2: fig.
I; pi. CI5: figs. 1-3; pi. C15: figs. 1-3, 5. 6. 8. 9.
Dendrophyllia boschmai cyathoheluxies [sic].—Song, 1991:136, pi. 1: figs.
7.8.
DIAGNOSIS.—Colonies robust and uniplanar, up to 9 cm in
height and width. Corallites primarily sympodially budded
from branch edges but in a very crowded manner, with
additional corallites on corallum faces. Cocnosteum costate and
nonepithecate. Corallites large, up to 9 x II mm in diameter.
Septa hexamerally arranged in 4 cycles in a typical Pourtales
Plan, larger corallites having some pairs of S5. Inner edges of
S1 smooth, those of S2_5 dentate. Columella well developed
and spongy. Corallum white; polyps red with yellow tentacles.
DISCUSSION.—No specimens of this species were examined
and thus the diagnosis above is taken from the literature. D.
boschmai differs from the other sympodially branched species
known from Japan, D florulenta, in having larger corallites, a
more robust corallum, and a much denser budding arrangement.
MATERIAL EXAMINED.—None.
TYPES.—The holotype of D. boschmai (= D. japonica of
Van der Horst, 1922) is deposited at the ZMA (Coel. 5451)
(Van Soest, 1979). Type Locality: "Japan," depth unknown.
The holotype of D. boschmai cyatheloides is deposited at the
TIUS (57443). Type Locality: Off Kowa, Mic-ken, Honshu,
depth unknown.
DISTRIBUTION.—Known only from the Japanese region:
Sagami and Suruga Bays, Honshu; off Kii Peninsula, Honshu;
off Shikoku; south of Cheju Do, South Korea; 40-165 m.
Dendrophyllia florulenta Akock, 1902
PLATE 39