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Theca marked by thin (0.05 mm wide), shallow, vertical striae
that delimit wide (0.5-0.6 mm), flat costae.
Septa hexamerally arranged in 4 cycles according to the
formula: S1_2>S3>S4, 4 pairs of S4 often lacking resulting in
40 septa. S^g about one-third LCD in width, their vertical,
slightly sinuous lower, inner edges fusing to the columella. S3
unflanked by S4 only about one-quarter width of an S, and
have finely serrate inner edges; however, if S3 are flanked by a
pair of S4, they are about three-quaters width of an S1 and have
smooth inner edges that fuse with the columella. S4 rudimentary,
only about 0.3-0.4 mm wide, and have finely serrate inner
edges. All septa relatively thin (S^_2 about 0.2 mm thick) and
widely spaced, i.e., 0.5-0.6 mm from one another. Fossa deep
and elongate, containing a well-developed, elongate trabecular
columella that occupies the medial third of fossa.
DISCUSSION.—Placotrochides is very similar to
Truncatoflabellum, but can be distinguished by its more robust
columella, and by having virtually parallel thecal edges and
faces. Only one other species is known in the genus, P. frustum
Cairns, 1979 (nom. correct), which is known only from the
Atlantic. It is distinguished by its smaller corallum size and
fewer (^26) number of septa.
MATERIAL EXAMINED.—Ate* Records: TM (KT7911,
OT4), 5,USNM 92748; TM (KT9202, YT1), 2, USNM 92749,
1, ORI. Previous Records: Holotype of P. scaphula; Philippine
specimens (Cairns, 1989a).
TYPES.—The holotype of P. scaphula is deposited at the
ZMA (Coel. 1094). Type Locality: Sihoga-2\2: 5°54.5'S,
120°19.2'E (Flores Sea), 462 m.
DISTRIBUTION.—Japan: Suruga Bay, Honshu; Osumi Shoto,
northern Ryukyu Islands (first records for Japan); 80-457 m.
Elsewhere: Philippines, Flores Sea; southwest Indian Ocean;
462-1628 m.
DIAGNOSIS.—See Part 1.
Javania Duncan, 1876
Javania insignis Duncan, 1876
PLATE 34/-*
Javania insignis Duncan, 1876:435. pi. 39: figs. 11-13.—Zibrowius, 1974b:
8-9. pi. 1: figs. 1-6.—Cairns, 1989a:77-78, pi. 40: figs. d,e,g,h.j,k
[synonymy].—Caims and Keller, 1993:272.
Flabellum weberi Alcock, 1902a: 107.
Desmophyllum cf. insigne.—Yabe and Eguchi, 1942b: 115, pi. 9: figs. 5, 6.
Desmophyllum insignis.—Eguchi. 1965:290, 2 figs.; 1968:C41-42, pi. C9:
figs. 4-9.—Song. 1982:136, pi. 2: figs. 5, 6; 1988:27-28, pi. 3: figs. 9-11;
1991:134.
DESCRIPTION.—Corallum ceratoid and robust, with thick
theca and septa, and a massive, stereome-reinforced pedicel
24%-55% of GCD. Largest Japanese specimen examined
(ZMC: Fukue Jima) 32.7 x 24.9 mm in calicular diameter and
56.2 mm in height, with a pedicel 7.8 mm in diameter, although
a slightly larger specimen was reported by Cairns (1984) from
Christmas Island. Theca milky white and smooth, sometimes
porcellaneous.
SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
Septa hexamerally arranged in 5 complete cycles according
to the formula: S1_2>S3>S4>S5 (96 septa). S^ exsert as much
as 4 mm and have quite thick, straight, vertical inner edges that
disappear into the fossa. S3 exsert as much as 2.5 mm, about
four-fifths width of an S^ and have relatively thin inner edges.
S4_5 nonexsert, the S4 only about one-third width of an S3, the
S5 about half width of an S4. S1 _2 often so thick that their
adjacent S5 are fused to them. Fossa very deep and elongate; no
columella visible.
DISCUSSION.—I (Cairns, 1989a:77) previously cited and
figured Alcock's "manuscript type" of Flabellum weberi as a
specimen of/, insignis. Because F. weberi was not reported by
Alcock (1902c) in his Siboga report, I assumed that he had
chosen not to describe this species. But since 1989, I have
become aware of two obscure papers published by Alcock
(1902a,b) in July, 1902, one month before his Siboga report
was published, in which he briefly described most of the
Siboga new species. However, for some reason, eight species
described in those earlier papers (Alcock, 1902a,b) were not
redescribed in the later Siboga report (Alcock, 1902c), F.
weberi being one of them. Thus, F. weberi is a legitimate junior
synonym of/, insignis, not merely a manuscript type.
Javania insignis is compared to /. borealis in the account of
that species.
MATERIAL EXAMINED.—New Records: TM (KT7811,
OT6-2), 1, USNM 92744; TM (KT9015, CB1-2), 1, USNM
92745; off Jogashima, Sagami Bay, 1, USNM 92746, 1, ORI;
off Misaki, Sagami Bay, 1,ZMC; 32°26'N, 128°37'E,249m, 1,
ZMC. Previous Records: Holotype of F. weberi, ZMA;
Philippine and Japanese specimens (Caims, 1989a).
TYPES.—The holotype of/, insignis is deposited at the BM
(1973.2.20.1). Type Locality: 34°13TST, 136°13'E (off Owase,
Mie-ken, Honshu), 88 m.
The holotype of F. weberi is deposited at the ZMA (Coel.
1232). Type Locality: Siboga-3\0: 8°3O'S, 119°07.5'E
(Flores Sea), 73 m.
DISTRIBUTION.—Japan: Sagami and Suruga Bays, Honshu;
off Owase, Mie-ken, Honshu; East China Sea off Fukue Jima;
Eastern Channel, Korea Strait, off southwestern Honshu;
southern Cheju Do, Korea; 46-249 m. Elsewhere: Widespread,
including Hawaiian Islands, Philippines, Indonesia,
Red Sea, and southwest Indian Ocean; 73-825 m.
Javania cailleti (Duchassaing and Michelotti, 1864)
ACCOUNT.—See Part 1.
ACCOUNT.—See Part 1.
Javania borealis sp. nov.
Rhiwtrochus Milne Edwards and Haime, 1848a
DIAGNOSIS.—Corallum ceratoid to turbinate or compressed.
Transverse division absent; pedicel small and not reinforced
with stereome. Thecal spines absent; however, 2-20 slender