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Aloes and Lilies of Ethiopia and Eritrea

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Distribution <strong>and</strong><br />

classification<br />

Reproduction<br />

HYACINTHACEAE<br />

SCHIZOBASIS DIPCADI 183<br />

The family includes bulbous plants with leaves, <strong>of</strong>ten more<br />

or less spotted, filiform to lanceolate, in a basal rosette.<br />

The scape is glabrous <strong>and</strong> leafless. The inflorescence is<br />

usually a raceme or a spike, rarely extremely branched<br />

(as in Schizobasis). The flowers are white to greenish,<br />

yellowish, or purplish, <strong>of</strong>ten with a darker green or<br />

purple midrib. The tepals are free or fused into a shorter<br />

or longer tube. The ovaries include one to many ovules<br />

per locule. The fruit is most <strong>of</strong>ten a loculicidal capsule,<br />

that means it splits between the walls separating its three<br />

chambers, rarely it is septicidal, i.e. it splits along the<br />

separating walls. The seeds are black, subglobose or<br />

flattened, sometimes winged.<br />

Hyacinthaceae is related to the Alliaceae (the onion<br />

family), differing mainly in chemistry <strong>and</strong> inflorescence,<br />

which is umbel­like in the latter. The sulphur compounds<br />

giving the particular smell <strong>and</strong> taste <strong>of</strong> members <strong>of</strong> the<br />

onion family are lacking in the hyacinths. Amaryllidaceae<br />

is also related, sharing the features <strong>of</strong> a typical bulb, the<br />

basal leaf rosette <strong>and</strong> the glabrous scape, differing in<br />

the umbel-like inflorescence similar to the onions; <strong>and</strong><br />

further by the inferior ovary.<br />

The family is widely distributed in tropical to temperate<br />

areas world­wide. Its centres <strong>of</strong> diversity are in Southern<br />

Africa <strong>and</strong> in the Mediterranean area. Taxonomists<br />

today disagree on the generic delimitation; from 40 to<br />

67 genera are recently reported. On species number the<br />

agreement is better, <strong>and</strong> c. 900 species are accepted.<br />

On the Horn <strong>of</strong> Africa there are 7 genera: Schizobasis,<br />

Dipcadi, Drimiopsis, Ledebouria, Drimia, Albuca, <strong>and</strong><br />

Ornithogalum. All these genera have centres <strong>of</strong> variation<br />

in southern Africa. Dipcadi, Drimia, <strong>and</strong> Ornithogalum<br />

reach the Mediterranean region <strong>and</strong> Central Asia. The<br />

species from the Horn <strong>of</strong> African find their closest<br />

relatives in the south.<br />

Pollen <strong>and</strong> nectar are <strong>of</strong>fered as floral rewards. The nectar<br />

is produced in the septal walls <strong>of</strong> the ovary, <strong>and</strong> secreted

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