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Aloes and Lilies of Ethiopia and Eritrea

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312 DIOSCOREACEAE<br />

Dioscorea gillettii<br />

Description<br />

Habitat <strong>and</strong><br />

distribution<br />

widespread to Senegal in the west <strong>and</strong> to Transvaal<br />

<strong>and</strong> Angola in the south. The main flowering period in<br />

<strong>Ethiopia</strong> is from August to September.<br />

This species includes wild varieties with poisonous<br />

tubers <strong>and</strong> bulbils with alkaloids (dioscorine <strong>and</strong> dihydrocorti<br />

sone). The wild tubers are eaten in times <strong>of</strong> famine,<br />

only after repeated washing.<br />

4. Dioscorea gillettii Milne-Redhead<br />

The specific epithet ‘gilletii’ was given in honour <strong>of</strong> the<br />

English botanist, Jan Gillett, who had collected much<br />

in eastern <strong>and</strong> ortheastern Africa. One <strong>of</strong> his collections<br />

from the Kenya­side <strong>of</strong> Moyale was designated as a type<br />

<strong>of</strong> the species. The species was described in 1963 by<br />

Milne­Redhead.<br />

It differs from a similar species, D. bul bifera, by the<br />

stems growing only up to 2 m long, male flowers with<br />

3 stamens, <strong>and</strong> with wingless seeds. In contrast, D.<br />

bul bifera has 3–10 m long stems, male flowers with 6<br />

stamens <strong>and</strong> winged seeds.<br />

The plant is a small herbaceous twiner, less than 2 m high. Stems<br />

unarmed, glabrous. Tuber perennial, spreading horizontally just<br />

below the soil surface with fringing roots, 3.5–4.5 x2.5–3.2 cm.<br />

Leaves alternate, rarely sub­opposite; petiole 0.5–5 cm long; blade<br />

ovate­cordate, 1.5–9 × 1–7.5 cm, acute to acuminate at the apex,<br />

deeply cordate at the base. Male inflorescences: In the axils <strong>of</strong><br />

leaves, 1–14 cm long; with single pedicillate flowers or short 2–4<br />

flower cymules at each node. Male flowers: perianth spreading, 4–5<br />

mm wide; segments elliptic; stamens 3 <strong>and</strong> staminodes 3. Female<br />

inflorescences: one raceme per leaf axil, usually 2-flowered, rarely<br />

one or more. Female flower: perianth spreading, 5–6 mm wide;<br />

ovary glabrous. Capsule obovate to suborbicular, 2.2–2.7 × 2–3 cm,<br />

winged, with parallel nerves extending from the suture towards the<br />

axis, not reflexed. Seeds without a wing.<br />

The species grows in Acacia-Commiphora-Boswellia<br />

woodl<strong>and</strong> <strong>and</strong> wooded grassl<strong>and</strong> between 850 <strong>and</strong> 1650<br />

m in Sidamo <strong>and</strong> Bale floristic regions. It also occurs in<br />

Kenya. The main flowering period in <strong>Ethiopia</strong> is from<br />

April to June <strong>and</strong> also from October to November.

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