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Aloes and Lilies of Ethiopia and Eritrea

Aloes and Lilies of Ethiopia and Eritrea

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Habitat <strong>and</strong><br />

distribution<br />

Dioscorea<br />

dumetorum<br />

Description<br />

Habitat <strong>and</strong><br />

distribution<br />

DIOSCOREA 311<br />

perianth subglobose c. 1 mm long, densely pubescent; stamens 6.<br />

Female inflorescences: pendulous spikes 15–20 cm long, in the<br />

axils <strong>of</strong> leaves. Capsule 5–7 × 2.5 cm, velutinous, reflexed upwards.<br />

Seeds with a wing on the basal side only.<br />

The species grows in broad­leaved deciduous woodl<strong>and</strong><br />

with Adansonia digitata <strong>and</strong> Boswellia papyrifera,<br />

along river valleys between 900 <strong>and</strong> 1300 m in Tigray<br />

<strong>and</strong> Gonder floristic regions. It also occurs in Tanzania,<br />

Burundi, Zaire, Zambia, Malawi, <strong>and</strong> Zimbabwe. The<br />

main flowering period in <strong>Ethiopia</strong> is from August to<br />

September.<br />

3. Dioscorea dumetorum (Kunth) Pax<br />

The specific epithet ‘dumetorum’ refers to the thicket<br />

(dumetum) habitat in which the plant grows.<br />

The species was described as Helmia dumetorum by<br />

Kunth in 1850 from plants collected in Tigray floristic<br />

region by Schimper. It differs from the related species,<br />

D. cochleari-apiculata by the 3-nerved leaflets <strong>and</strong><br />

the glabrous perianth <strong>and</strong> mature capsule glabrous. In<br />

contrast, D. cochleari-apiculata has 3-nerved leaflets,<br />

<strong>and</strong> perianth <strong>and</strong> mature capsule pubescent.<br />

Prickly twiner or climber, stems growing 8–10 m long. Bulbils in<br />

the axils <strong>of</strong> leaves, prickly, 3–5 cm long, pubescent. Tubers renewed<br />

annually with several per plant; each tuber subglobose or cylindrical.<br />

Leaves alternate, 3­foliolate, pubescent above, tomentose below,<br />

rarely glabrescent; petiole 5–20 cm long, generally prickly; petioles,<br />

0.3–1 cm long; lateral leaflets obovate, 5–16 × 3–15 cm, acute to<br />

acuminate at the apex, cuneate or rounded at the base, conspicuously<br />

3-nerved from just above the base. Male inflorescences: axillary or<br />

terminal, much branched panicles <strong>of</strong> dense spikes; spikes short,<br />

0.5–1(–1.5) cm long, subsessile or with peduncle up to 0.3 cm<br />

long, cylindrical; bracts broadly ovate, adpressed to the perianth<br />

<strong>and</strong> partly concealing it, densely pubescent. Male flowers: perianth<br />

subglobose, glabrous, opening little; stamens 6 at least in the first<br />

flowers <strong>and</strong> 5 or fewer in successive ones. Female inflorescences:<br />

pendulous axillary spikes, 10–20 cm long. Female flowers: directed<br />

downwards, s<strong>of</strong>tly tomentose all over; ovary densely pubescent.<br />

Capsules directed upwards, 2.5–4.5 × 1.7–2.4 cm, glabrous to<br />

sparsely pubescent. Seeds with a wing on the basal side only.<br />

The species grows in Combretum-Termi nalia woodl<strong>and</strong><br />

<strong>and</strong> secondary thickets, <strong>and</strong> along river valleys between<br />

900 <strong>and</strong> 1525 m in Tigray, Gonder, Gojam, <strong>and</strong> Wellega<br />

floristic regions in <strong>Ethiopia</strong> <strong>and</strong> in <strong>Eritrea</strong>. It is also

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