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Aloes and Lilies of Ethiopia and Eritrea

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206 ALOACEAE<br />

Distribution <strong>and</strong><br />

classification<br />

Reproduction<br />

Use <strong>and</strong><br />

chemistry<br />

ALLIACEAE<br />

The family includes herbs with bulbs, bulb­like corms, or<br />

rhizomes. The underground parts usually produce a smell<br />

<strong>of</strong> garlic or onion when squashed. Leaves are produced<br />

in a basal rosette, or they are 2-ranked. The inflorescence<br />

is an umbellate or a globose head, subtended by 1–2<br />

membranous bracts. The flowers are commonly regular,<br />

the perianth varying from white to pink, blue, violet or<br />

purple, sometimes yellow. The 6 tepals are more or less<br />

equal. The stamens are attached to the base <strong>of</strong> the tepals.<br />

The filaments are flattened. The ovary is superior with<br />

nectar producing groves in the walls. The fruit is a capsule<br />

with few to many seeds, <strong>of</strong>ten somewhat triangular in<br />

cross section.<br />

The family Alliaceae is mainly distributed in the temperate<br />

regions, both in the southern <strong>and</strong> northern hemispheres.<br />

Only a few species reach the tropics. It is represented by<br />

c.13 genera <strong>and</strong> 700 species worldwide. In the Flora area,<br />

it is represented by two genera: Allium <strong>and</strong> Agapanthus.<br />

The genus Agapanthus, represented by A. praecox, is<br />

only known in cultivation in <strong>Ethiopia</strong> <strong>and</strong> <strong>Eritrea</strong>, <strong>and</strong> is<br />

not further treated here. The genus Allium is represented<br />

by indigenous <strong>and</strong> cultivated taxa.<br />

A large number <strong>of</strong> species with bulbs have vegetative<br />

propagation. Often some <strong>of</strong> the flowers are substituted by<br />

bulbils that function as vegetative dispersal units.<br />

One <strong>of</strong> the characteristic features <strong>of</strong> the family is the<br />

absence <strong>of</strong> alkaloids which are found in related families as<br />

Amaryllidaceae. Members <strong>of</strong> the genus Allium are known<br />

to contain sulphur­compounds which contribute to their<br />

particular smell <strong>and</strong> taste. They also include flavones <strong>and</strong><br />

flavonols. In fact, Allium species can be separated into<br />

three groups based on these compounds: those that have<br />

only flavones, those with only flavonols <strong>and</strong> those with<br />

both. Species <strong>of</strong> the genus have been in cultivation since<br />

the first periods <strong>of</strong> agriculture in the Old World. Onions,<br />

garlic, leek, <strong>and</strong> chives belong in the genus.

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