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sdu faculty of forestry journal special edition 2009 - Orman Fakültesi

sdu faculty of forestry journal special edition 2009 - Orman Fakültesi

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SDÜ ORMAN FAKÜLTESİ DERGİSİ<br />

Canker width was correlated to canker length, which was due to canker shape,<br />

since the majority <strong>of</strong> cankers were oval shaped. Regression analysis with a linear<br />

model (canker width = 14.6 + 0.44 × canker length) gave correlation coefficient <strong>of</strong><br />

0.87. Coefficient 0.44 tells that canker width measured usually about a half <strong>of</strong><br />

canker length.<br />

Possible entry points for E. parasitica are branch stubs and bark wounds<br />

(French, 1969). Great majority <strong>of</strong> cankers were traced to branch stubs as the<br />

avenue <strong>of</strong> entry. Only 3 out <strong>of</strong> 89 cankers were associated with bark wounds and<br />

all three wounds resulted from logging operations.<br />

3.3. Area and shape <strong>of</strong> canker on the trunk<br />

The largest Eutypella canker measured 1.7 m 2 , although the average was much<br />

lower, i.e. 0.5 m 2 (Table 2). The canker area varied a lot because it depended upon<br />

the canker age. The cankered area ranged from 4% to 90% <strong>of</strong> the trunk area with<br />

the canker, with an average <strong>of</strong> 48%.<br />

The area without bark also depended upon the canker age and thus the whole<br />

canker area. In order to determine if in fact there was a relationship between the<br />

canker area without bark and whole canker area, regression analysis using a linear<br />

model was carried out. The linear model described a relationship between canker<br />

area without bark and whole canker area. The equation <strong>of</strong> the model was: area<br />

without bark = –2.32 + 0.25 × whole canker area. The high correlation coefficient<br />

(0.83) showed a good linear relationship between the canker area without bark and<br />

whole canker area. The coefficient <strong>of</strong> the equation tells that the area without bark<br />

represented usually a quarter <strong>of</strong> the whole canker area when the canker was old<br />

enough to bark fall <strong>of</strong>f. The bark begins to fall <strong>of</strong>f a trunk due to high degree <strong>of</strong><br />

decay or/and high activity <strong>of</strong> bark beetles.<br />

Perithecia do not usually develop nearer than the fourth or fifth annual callus<br />

ring from the margin <strong>of</strong> the cankers (Davidson and Lorenz, 1938). Therefore,<br />

Davidson and Lorenz (1938) stated that perithecia are confined to only a small<br />

central portion <strong>of</strong> large cankers. During this study, their hypothesis about the area<br />

size covered with perithecia was tested. The part <strong>of</strong> their hypothesis was rejected. It<br />

was confirmed that perithecia are usually located in the central portion <strong>of</strong> the<br />

canker but this area is not small. The total area <strong>of</strong> bark covered with perithecia<br />

could exceed 68% <strong>of</strong> the whole canker area. An average <strong>of</strong> 32% (±7% at 95%<br />

confidence level) <strong>of</strong> the canker area was covered with perithecia. Further analysis<br />

was performed and it was found that the correlation between the total area <strong>of</strong><br />

perithecia and the area <strong>of</strong> the whole canker was very good. Simple regression<br />

analysis produced a correlation coefficient <strong>of</strong> 0.94. A simple regression was made<br />

using a linear model: total area <strong>of</strong> perithecia = –3.93 + 0.42 × whole canker area.<br />

Three different densities <strong>of</strong> perithecia were delineated. The high densities <strong>of</strong><br />

perithecia were usually located nearer to the canker centre while low densities were<br />

located nearer to canker margin. It should be noted that a middle density <strong>of</strong><br />

perithecia could hardly be distinguished from high density. Therefore, some<br />

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