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A. Status of the Spectacled Eider - U.S. Fish and Wildlife Service

A. Status of the Spectacled Eider - U.S. Fish and Wildlife Service

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APPENDIX II<br />

TECHNICAL DETAILS OF THE BAYSIAN ANALYSIS<br />

CALCULATION OF POSTERIOR DISTRIBUTIONS<br />

Estimation <strong>of</strong> <strong>the</strong> amount <strong>of</strong> risk to which <strong>Spectacled</strong> <strong>Eider</strong>s are exposed must be based<br />

primarily on an analysis <strong>of</strong> <strong>the</strong> rate <strong>of</strong> change <strong>of</strong>population abundance where abundance is<br />

estimated by various survey techniques. For <strong>the</strong> <strong>Spectacled</strong> <strong>Eider</strong> three time series <strong>of</strong><br />

abundance estimates are analyzed: <strong>the</strong> North American breeding bird survey (number <strong>of</strong><br />

estimates (n) = 39), aerial surveys <strong>of</strong> <strong>the</strong> YKD coast (n = 8), <strong>and</strong> ground plot surveys where<br />

r<strong>and</strong>om ground plots stratified by habitat type are exhaustively surveyed for nests (n = 10).<br />

For each survey we would like to know <strong>the</strong> probability <strong>of</strong> obtaining <strong>the</strong>se data given various<br />

hypo<strong>the</strong>ses concerning <strong>the</strong> unknowns: population growth rate (r--<strong>the</strong> slope estimate <strong>of</strong> a<br />

regression <strong>of</strong> population size against time) <strong>and</strong> variance in <strong>the</strong> estimate <strong>of</strong> population growth<br />

rate (<strong>the</strong> st<strong>and</strong>ard error <strong>of</strong><strong>the</strong> estimate). Given a probability distribution for population<br />

growth rate for our particular set <strong>of</strong> data we can directly answer questions about <strong>the</strong> probability<br />

that <strong>the</strong> population is stable or growing. The range <strong>of</strong> hypo<strong>the</strong>ses tested constitute <strong>the</strong> prior<br />

distribution <strong>and</strong> <strong>the</strong> resultant distribution, which is conditional on <strong>the</strong> data, is called <strong>the</strong><br />

posterior distribution.<br />

Our primary interest is in estimating <strong>the</strong> population growth rate from observed data, which are<br />

abundance index estimates with associated precision estimates. We assume an exponential model<br />

<strong>of</strong>population growth (equation 1).<br />

= N 0 e(ree) t (1) (2.)<br />

where N = number <strong>of</strong>breeding pairs, t = time (years), N0 = initial number <strong>of</strong>breeding pairs, r =<br />

population growth rate <strong>and</strong> E~ is Gaussian distribution symbolized as G (~, 5r2), where ~ = 0 <strong>and</strong> 5r<br />

is <strong>the</strong> st<strong>and</strong>ard deviation <strong>of</strong>r. When s~ = 0 <strong>the</strong> model is deterministic. When ~r > 0 <strong>the</strong>n <strong>the</strong><br />

model is stochastic with annual growth rate drawn from a distribution. The parameter <strong>of</strong>interest.<br />

for classification decisions is r. Parameters are estimated by fitting <strong>the</strong> model with <strong>the</strong> available<br />

time-series <strong>of</strong>abundance estimates using Bayesian methods. This is analogous to a weighted nonlinear<br />

regression using classical statistical methods (essentially because <strong>the</strong> contribution <strong>of</strong>each<br />

abundance estimate to <strong>the</strong> estimation is weighted by <strong>the</strong> inverse <strong>of</strong>its estimated precision or CV).<br />

Due to <strong>the</strong> nature <strong>of</strong><strong>the</strong> available data set, several o<strong>the</strong>r parameters have to be defined <strong>and</strong><br />

estimated. The abundance estimates <strong>and</strong> <strong>the</strong>ir associated coefficients <strong>of</strong>variation, CV (which<br />

were estimated from <strong>the</strong> sampling design for each survey), represent <strong>the</strong> observed data in this<br />

analysis. Because <strong>the</strong> number <strong>of</strong> eiders that breed <strong>and</strong> are thus available to be counted may vary<br />

through time (more than explained by trends in r), <strong>the</strong> estimated CV may not account for all <strong>of</strong><strong>the</strong><br />

Appendix II- Page 1

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