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A. Status of the Spectacled Eider - U.S. Fish and Wildlife Service

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S 1(t) = m1 CV(N(t)) N(t) (2.0)<br />

where<br />

= <strong>the</strong> CV multiplier for survey I,<br />

CV(N1(t)) = <strong>the</strong> estimated survey precision (CV) <strong>of</strong>abundance estimate 141(t).<br />

Note that parameter values that maximize equation 4 are <strong>the</strong> maximum likelihood estimates, a<br />

common point estimator in frequentist statistics. In Bayesian statistics, ra<strong>the</strong>r than maximizing<br />

equation 4 we need to integrate <strong>the</strong> product <strong>of</strong>it <strong>and</strong> <strong>the</strong> prior distribution <strong>of</strong><strong>the</strong> parameters<br />

(defined below).<br />

Until fairly recently, most Bayesian analyses were restricted to cases where <strong>the</strong> prior distribution<br />

could be chosen so that it was “conjugate” to <strong>the</strong> likelihood distribution, resulting in <strong>the</strong>ir product<br />

being a distribution <strong>of</strong>a known form, which was integratable by analytic methods. Advances in<br />

computing power <strong>and</strong> numerical <strong>and</strong> Monte Carlo integration methods have removed this<br />

restriction. We use <strong>the</strong> Sampling-Importance-Resampling routine <strong>of</strong>Rubin (1988), which Smith<br />

<strong>and</strong> Gelf<strong>and</strong> (1992) advocate as a particularly useful <strong>and</strong> simple integration technique for<br />

Bayesian statistics. In this method, values for <strong>the</strong> parameters are r<strong>and</strong>omly selected from <strong>the</strong>ir<br />

joint prior distribution to form a sample set 0~. The likelihood <strong>of</strong><strong>the</strong> data given this particular O~<br />

is calculated <strong>and</strong> stored. This is repeated, generating n1 O,’s with associated likelihoods. These n1<br />

O,’s are <strong>the</strong>n re-sampled n2 times with replacement, with probability equal to weight q~, where<br />

L(011x) (12.)<br />

j~1 L(831x)<br />

Rubin (1988) showed that this generates a r<strong>and</strong>om sample from <strong>the</strong> joint posterior distribution <strong>of</strong><br />

size n2. The resampling with replacement from <strong>the</strong> n1 O,’s with weight CL makes this process<br />

analogous to a weighted bootstrap procedure. We set values for n1 <strong>and</strong> n2 to yield smooth<br />

posterior distributions, which depended on <strong>the</strong> number <strong>of</strong>parameters estimated. For single survey<br />

analyses, n1 = 300,000, n2 = 5,000. For analyses <strong>of</strong>all three surveys, n1 = 8,000,000, n2 = 10,000.<br />

For analyses <strong>of</strong>all surveys with environmental stochasticity (equation 1), n1 = 25,000,000, n2 =<br />

10,000. With <strong>the</strong> exception <strong>of</strong>initial population size, prior distributions were Uniform<br />

distributions that were iteratively set to encompass <strong>the</strong> values <strong>of</strong> <strong>the</strong> posterior distribution for each<br />

parameter.<br />

For <strong>the</strong> full analysis using <strong>the</strong> deterministic population model (equation 1) <strong>and</strong> all three data sets,<br />

values were r<strong>and</strong>omly drawn from <strong>the</strong>ir prior distributions for <strong>the</strong> seven parameters (r, N0 ma, 1%,<br />

a0~ <strong>and</strong> ar)) to form a 01. Those parameter values were <strong>the</strong>n used to project a model<br />

population trajectory using equation 1, forming a series <strong>of</strong>model population sizes N1, N2, . .N~.<br />

The likelihood <strong>of</strong>that population trajectory given <strong>the</strong> data was <strong>the</strong>n calculated using equation 9,<br />

<strong>and</strong> <strong>the</strong> 0, ‘s <strong>and</strong> associated likelihood value were stored. This was repeated n1 times to form <strong>the</strong><br />

Appendix II- Page 4

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