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260<br />

B. Baur and S. Schmidlin<br />

(Grand Canal d’Alsace). During channelisation, flood control dams were<br />

built, stone groynes were constructed to strengthen the channel, and some<br />

parts of the river bank were reinforced by stones.<br />

Since Roman times, the Rhine has been a navigable waterway, carrying<br />

travellers and goods deep inland. As the river became more important as<br />

transport route, it was channelled even more to increase its discharge and<br />

maintain its depth. In modern times, cargo shipping on the Rhine is possible<br />

from Rotterdam (North Sea) to Rheinfelden, 20 km upstream of Basel. The<br />

importance of international shipping increased further with the construction<br />

of river-connecting canals. The Rhine-Main-Danube Canal connects the<br />

Rhine via the Danube with the Black Sea which, in turn, is connected by canals<br />

and rivers to the Caspian Sea (see Chap. 5). Another navigation route to the<br />

Black Sea and Caspian Sea is the German Mittelland-Elbe-Vistula-Pripyat-<br />

Bug-Dnieper canal system.<br />

Parallel to the channelisation, the floodplain of the Rhine has been extensively<br />

modified to extend agricultural and industrial areas and settlements.<br />

Nowadays, the Rhine is a completely man-manipulated river, more intensively<br />

used than ever before (Tittizer and Krebs 1996). Besides its function as transportation<br />

route, it provides water for communities and industry, is used to<br />

generate hydroelectric power, provides cooling water and a means of effluent<br />

transport, and is increasingly a focus for recreation. Despite profound alterations<br />

of river characteristics, the river still has a large (albeit not unlimited)<br />

self-cleaning capacity, and natural and semi-natural banks and areas of the<br />

floodplain, with abandoned meanders, brooks, sand and gravel pits, and remnants<br />

of riparian forest still harbour an extraordinarily high diversity of<br />

plants and animals, and are therefore of high conservation value (e.g. LfU<br />

2000; Baur et al. 2002).<br />

15.3 Native Biodiversity and Invasion History<br />

Faunal diversity decreased dramatically in the river Rhine between 1900 and<br />

1970 (Kinzelbach 1972; Van den Brink et al. 1990; Streit 1992). For example,<br />

species richness of selected groups of macroinvertebrates in the Dutch section<br />

of the Rhine declined from 83 species in 1900 to 43 species in 1940 and to<br />

41 species in 1981/1987 (Van den Brink et al. 1990; Den Hartog et al. 1992).<br />

Omitting the non-native species arriving in the 20th century, however, the<br />

total number of species for 1940 would be 40, and only 27 for 1981/1987. Schöll<br />

(2002) presented a list of 21 typical riverine macroinvertebrate species (seven<br />

mayflies, 10 stoneflies and four caddis flies) occurring in the German part of<br />

the Rhine in 1900 – none were found in the river between 1960 and 2000. Most<br />

probably, these specialized benthic species went extinct in the river Rhine.<br />

However, the actual causes of extinction are unknown. In the Rhine near

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