Type-data.-Female holotype from Stowers<strong>Cave</strong>, 24 miles W Kerrville, Kerr County, Texas, 3May 1969 (Roger Bartholomew) (AMNH).Distribution.-Known only from Stowers <strong>Cave</strong>.Other records: 20 March 1965 (J. Reddell), 5 immature;25 March 1972 (S. Wiley, T. Mollhagen), 1immatureCicurina pastura, new speciesFigs. 123-124Diagnosis.-Eyeless troglobite from Water PondPasture <strong>Cave</strong>, Kerr County, Texas; oval spermathecumwith rounded sac set in oblique position,with tips nearly touching; index coil thick canal inclinedoutward; connecting canal loosely enclosingspermathecal parts. Male unknown.Etymology.-Specific name from Latin pasture,pasture.Description.-Female holotype: Length 4 mm.Carapace 2 mm long, 1.4 mm wide. Abdomen 2 mmlong, 1.4 mm wide. Chelicerae: retromargin with 5teeth. Leg lengths: first femur 1.8 mm, fourth femur2 mm; first patella-tibia 2.2 mm, fourth patella-tibia2.5 mm. Leg spines: first tibia 2-2-0, fourth tibia2-2-2.Type-data.-Female bolotype from Water PondPasture <strong>Cave</strong>, Kerr County, Texas, 16 October 1976(D. Pate, R. Fieseler, C. Yates) (AMNH).Cicurina machete, new speciesFigs. 125-126Diagnosis.-Eyeless troglobite of Whiteface<strong>Cave</strong>, San Saba County, Texas; oval sperrnathecumwith twisted sac; index coil heavy procurvedblade-like canal crossing sac; connecting canalheavy loop around sperrnathecal parts. Male unknown.Etymology.-Specific name <strong>for</strong> Spanish machete,cutlass, used in apposition.Description.-Female holotype: Length 4.6 mm.Carapace 2.4 mm long, 1.6 mm wide. Abdomen 2.2mm long, 1.4 mm wide. Chelicerae: retromargin offang with 8 teeth. Leg lengths: first femur 2.4 mm,fourth femur 2.8 mm; first patella-tibia 3.2 mm,fourth patella-tibia 3.4 mm. Leg spines: first tibia2-2-2, fourth tibia 2-2-0.Type-data.-Female holotype and 4 immaturefrom Whiteface <strong>Cave</strong>, 30 mi. S Richland Springs, 9February 1964 (J. Reddell, D. McKenzie, K. Garrett)(AMNH).Cicurina sansaba, new speciesFigs. 127-128, 153-154Diagnosis.-Eyeless troglobite of Gorman <strong>Cave</strong>,San Saba County, Texas; elongate sperrnathecumwith broad rounded sac; index coil with transversecanal across sac; heavy connecting canal <strong>for</strong>mingloose loop around sperrnathecal parts. Male palpus:Figs. 153-154.Etymology.-Specific name <strong>for</strong> San SabaCounty.Description.-Female holotype: Length 3.65mm. Carapace 1.65 mm long, 1.15 mm wide. Abdomen3.5 mm long, 2.2 mm wide. Chelicerae:retromargin of fang with 6 teeth. Leg lengths: firstfemur 1.85 mm, fourth femur 1.95 mm; firstpatella-tibia 1.3 mm, fourth patella-tibia 3.25 mm.Ventral leg spines: first tibia 2-2-1, fourth tibia2-2-2.Male: Length 2.7 mm, specimen dried andshriveled. Male palpus (Figs. 153-154): tegulumwith three tubules; tarsal process about half width oftarsus; coil of conductor about length of basal process.Type-data.-Female holotype and 3 immaturefrom Gorman <strong>Cave</strong>, 6 mi. SW of Bend, San SabaCounty, Texas, 15 March 1963 (J. Reddell, D.McKenzie) (AMNH).Distribution.-Known only from Gorman <strong>Cave</strong>.Other records: 12 June 1978 (J. Reddell), 1 male, 2females, 2 immature, from Beyond Breakdown; 20October 1962 (J. Reddell), 1 immature.Cicurina venefica, new speciesFigs. 129-130Diagnosis.-Eyeless troglobite of Wizard's Well,Terrell County, Texas; elongate spermathecum withrounded sac, set in oblique position with tips nearlytouching; index coil large canal in inclined position;connecting canal <strong>for</strong>ming wide coil around spermathecalparts. Male unknown.Etymology.-Specific name from Latin venefica,a witch.Description.-Female holotype: Length 4.6 mm.Carapace 2.3 mm long, 1.6 mm wide. Chelicerae:retromargin of fang with 6 teeth. Leg lengths: firstfemur 2.2 mm, fourth femur 2.6 mm; firstpatella-tibia 3 mm, fourth patella-tibia 3.3 mm. Legspines: first and fourth tibiae with 2-2-0.Type-data.-Female holotype from Wizard'sWell, Terrell County, Texas, 12-13 February 1983(Eric Short, Randy Waters) (AMNH).114
Cicurina caverna, new speciesFigs. 131-132Diagnosis.-Eyeless troglobite of Flemming'sBat <strong>Cave</strong>, Kimble County, Texas; elongate ovalspermathecum with rounded sac; index coil procurvedcanal crowning sac; slender connecting canalfonning loose loop around sperrnathecal parts. Maleunknown.Etymology.-Specific name from Latin caverna,a cavern.Description.-Female holotype: Length 3 rnm.Carapace 2 mm long, 1.4 mm wide. Abdomen 2 rnmlong, 1.2 mm wide. Chelicerae: retromargin of fangwith 5 teeth. Leg lengths: first femur 1.6 mm,fourth femur 1.8 mm; first patella-tibia 1.2 rnm,fourth patella-tibia 1.4 mm. Leg spines: first andfourth tibiae 2-2-2.Type-data.-Female holotype and 5 immaturefrom Flemming's Bat <strong>Cave</strong>, 4 mi. N Telegraph,Kimble County, Texas, 21 February 1964 (W.H.Russell) (AMNH).Cicurina porteri, new speciesFigs. 133-134Diagnosis.-Eyeless troglobite of Oriente MilestoneMolasses Bat <strong>Cave</strong>, Val Verde County, Texas;elongate sperrnathecum with rounded sac; index coilheavy procurved canal across sac; connecting canalloosely enclosing spermathecal parts. Male unknown.Etymology.-Named <strong>for</strong> John Porter, student ofcaves.Description.-Female holotype: Length 4.4 rnm.Carapace 2.4 mm long, 1.8 mm wide. Abdomen 2mm long, 2.4 mm wide. Chelicerae: retromargin offang with 5 or 6 teeth. Leg lengths: first femur 2.2mm, fourth femur 2.4 mm; first patella-tibia 2.6mm, fourth patella-tibia 2.7 mm. Leg spines: firsttibia 2-2-0, fourth tibia 2-2-2.Type-data.-Female holotype and 1 immaturefrom Oriente Milestone Molasses Bat <strong>Cave</strong>, about20 mi. NE Del Rio, Val Verde County, Texas, 25January 1964 (1. Reddell, D. McKenzie, J. Porter),under rock beyond bat room (AMNH).Cicurina sheari, new speciesFigs. 135-136Diagnosis.-Eyeless troglobite of Ramsey Bat<strong>Cave</strong>, Real County, Texas; small coiled spermathecumand rounded sac in oblique position; indexcoil heavy recurved canal <strong>for</strong>ming invertedV-shaped figure; connecting canal fonning wideloop around sperrnathecal parts. Male unknown.Etymology.-Named <strong>for</strong> William A. Shear, studentof spider behavior and evolution.Description.-Female holotype: Length 4.7 mm.Carapace 2.2 mm long, 1.8 mm wide. Abdomen 2.5mm long, 1.8 mm wide. Chelicerae: retromarginwith 4 large teeth on left chelicera and 6 in scatteredrow on right chelicera. Leg lengths: first femur 2.2mm, fourth femur 2.5 mm; first patella-tibia 2.6mm, fourth patella-tibia 2.8 mm. Leg spines: firsttibia 2-2-1, fourth tibia 2-2-2.Type-data.-Female holotype from Ramsey Bat<strong>Cave</strong>, Real County, Texas, 2 October 1976 (D.Pate, R. Hemperly, K. Heuss) (AMNH).Cicurina suttoni, new speciesFigs. 137-138, 151-152Diagnosis.-Eyeless troglobite of Felton <strong>Cave</strong>,Sutton County, Texas; elongate sperrnathecum withbroadly rounded sac; index coil transverse inwardlyturned canal around sac; connecting canal fonning aloose loop around spermathecal parts. Male (Figs.151-152).Etymology.-Specific name <strong>for</strong> Sutton County.Description.-Female holotype: Length 5.8 mm.Carapace 2.6 mm long, 2 mm wide. Abdomen 3.2mm long, 2 mm wide. Chelicerae: retromargin offang with 6 teeth. Leg lengths: first femur 2.4 mm,fourth femur 2.5 mm; first patella-tibia 2.4 mm,fourth patella-tibia 2.5 mm. Leg spines: first tibia2-2-0, fourth tibia 2-2-2.Male: Length 5.4 mm. Carapace 2.6 mm long, 2mm wide. Abdomen 2.8 mm long, 2 mm wide.Chelicerae: retromargin of fang with 6 teeth. Leglengths: first femur 2 mm, fourth femur 2.4 mm;first patella-tibia 2.6 mm, fourth patella-tibia 2.8mm. Male palpus (Figs. 151-152).Type-data.-Female holotype from Felton <strong>Cave</strong>,15 mi. SW Sonora, Sutton County, Texas, 4 July1964 (J. Reddell), in rotting root 2000 feet from entrance(AMNH).Distribution.-Known only from Felton <strong>Cave</strong>.Other records: 4 JulY 1964 (J. Reddell), 6 immaturefrom near Bat Room, 600 ft. from entrance; 14 October1928 (O.G. Babcock), 940 feet from entrance,1 male.Cicurina mckenziei, new speciesFigs. 139-140Diagnosis.-Eyeless troglobite of Fog Fissure,Bandera County, Texas; rounded spermathecum115
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PREFACEThe present volume is the se
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TABLE OF CONTENTSHOLSINGER, JOHN R.
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the state of Coahuila in northern M
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Fig. 2.-Anesia welboumi, new specie
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\\. \ - -' ..........---~\ I// --..
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Mexiweckelia hardeni, new speciesFi
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2 sets of 1 or 2 setae each; dactyl
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AFig. 7.-Holsingerius smaragdinus,
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have as many setae on the inner pla
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Antenna 1 about 33 % length of body
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Of biogeographic interest for the h
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Bowman, T .E. 1992. Two subterranea
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A note by Scott Harden that accompa
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~~..~ ~~ I))\\ "-,'.=bFig. 3.-Speoc
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unarmed, except in pleopod 2, which
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Cokendolpher, LC., and l.R. Reddell
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lobes. The larger setae vary greatl
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the relationships of the order and
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zomids their absence is considered
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some species could be either split,
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have small pores over the surface o
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inhabits tropical deciduous forest
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huitvnolotitlensis from A. stygius
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8-10, figs. 5-7; Rowland, 1973c:136
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(in row) and one pair large posteri
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(0.34); tarsus 0.64 (0.58); total 4
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Abdomen: Tergite I with two pairs a
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setae near posterior margin. stemit
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setae, and ten ventral setae. Stemi
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1,980 m elev., 26 Dec. 1986 (T. Tre
- Page 73 and 74: asitarsal-tarsal proportions: 15:4:
- Page 75 and 76: and one pair setae at base of proce
- Page 77 and 78: Cephalothorax: Propeltidium 1.66 mm
- Page 79 and 80: Male adult unknown.Immature paratyp
- Page 81 and 82: IV: trochanter 1.20 (1.10); femur 3
- Page 83 and 84: Cokendolpher, 1.C. 1981. The order
- Page 85 and 86: Gertsch, W.J. 1992. Distribution pa
- Page 87 and 88: same families and genera, but also
- Page 89 and 90: species from the United States and
- Page 91 and 92: the posterior median pair short and
- Page 93 and 94: great many species of North America
- Page 95 and 96: Key to the Eyed Females1. Eight eye
- Page 97 and 98: 39111012Figs. 1-12.-Ventral and dor
- Page 99 and 100: 1419 23Figs. 13-24.-Ventral and dor
- Page 101 and 102: 27 293334Figs. 25-36.-Ventral and d
- Page 103 and 104: Figs. 37-48.-Ventral and retrolater
- Page 105 and 106: Cicurina blanco, new speciesFigs. 7
- Page 107 and 108: Type-data.-Female holotype from ins
- Page 109 and 110: Description.-Female holotype: Lengt
- Page 111 and 112: Description.-Female holotype: Lengt
- Page 113 and 114: lengths: first femur 2 rom, fourth
- Page 115 and 116: Cicurina pablo, new speciesFigs. 10
- Page 117 and 118: Description.-Female holotype: Lengt
- Page 119 and 120: canal in nearly vertical posItion;
- Page 121 and 122: Cicurina vespera, new speciesFigs.
- Page 123: procurved canal across sac; connect
- Page 127 and 128: with sac of similar size set in obl
- Page 129 and 130: February 1964 (J. Reddell, D. McKen
- Page 131 and 132: County: Diamond Cave, 16 August 196
- Page 133 and 134: Roth, V.D. 1992. A new and first tr
- Page 135 and 136: LITERATURE CITEDBarr, T.C. 1963. Ec
- Page 137 and 138: Muchmore, W.B. 1992. Cavernicolous
- Page 139 and 140: Species of Aphrastochthonius are kn
- Page 141 and 142: FAMILY NEOBISIIDAE CHAMBERLINGenus
- Page 143 and 144: trochanter 2.5 (2.6), femur 5.55 (5
- Page 145 and 146: ottom of entrance pit, Ogle Cave (O
- Page 147 and 148: tactile seta on tibia and basitarsu
- Page 149 and 150: Missouri may be conspecific (unpubl
- Page 151 and 152: Trichobothriotaxy of chela generall
- Page 153 and 154: Female (figures given first for all
- Page 155 and 156: transverse furrows; eyespots not ev
- Page 157 and 158: Reddell and W. Russell); I female f
- Page 159 and 160: and Acuminochernes, along with the
- Page 161 and 162: Chamberlin, J.C. 1946. The genera a
- Page 163: (0.36); chela (without pedicel) 2.0
- Page 167 and 168: although not recently studied is no
- Page 169 and 170: and stylar outgrowths, present in s
- Page 171 and 172: in Phalangodes (et al.) is thick an
- Page 173 and 174: pairs) are found in six species: T.
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elated species IS probably best int
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legs than expected (2.6-3.2). This
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~ 3.3 are synapomorphic); all trogl
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TAXONOMYTEXELLA Goodnight and Goodn
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14. BK absent (Figs. 177, 180). SA
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Figs. 8-11.-Texella bijUrcata (Brig
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Description.-Total body length, 1.5
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male examined closely has fewer set
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Figs. 26-29.-Texella kokoweej, new
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apical region which loses the apica
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Figs. 38-41.-Texella shoshone, new
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Distribution.-Known only from the t
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Figs. 52-55.-Texella brevistyla, ne
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Texellajungi, new speciesFigs. 60-7
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Figs. 66-69.-Texellajungi, new spec
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than S; SA with laterobasal carina
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~81//J/'/ ;'?/ ~~.........--~~I, II
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cylindrical, retina and cornea abse
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Figs. 93-96.-Texella cokendolpheri,
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SA with well developed prong and re
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Figs. 105-108.-Texelia mulaiki Good
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and McCarty Caves, which are known
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Color orange. Body of medium rugosi
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Notes.-The type locality was errone
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Figs. 128-131.-Taella reyesi, new s
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Figs. 136-139.-Texella reyesi, new
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Figs. 144-147.-Texella reyesi, new
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Figs. 152-155.-Texella reyesi, new
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1989 (W. Elliott, J. Reddell, and M
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Table 3.-Continued.# locality sex S
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mesoapical; patella, 2 mesal; tibia
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Figs. 162-165.-Texella gmbbsi, new
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Figs. 166-169.-Texella diplospina,
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Figs. 174-177.-Texella renkesae, ma
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Figs. 178-18\.-Teulla spinoperca, n
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Distribution.-Known only from Fayet
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Figs. 190-193.-Texellafendi, new sp
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CLASSIFICAnONTexellabifurcata group
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Chandler, D.S. 1992. The Pselaphida
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Key to Species1. Abdominal segments
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stemite VI slightly impressed at ba
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vertexal carinae, and the laterally
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is associated with rotten woods (Ch
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small ventral carina near base, pro
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Grigarick, A.A., and R.O. Schuster.
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Cicurifla (Cicurella) holsiflgeri G