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Key to Genera of Cavernicolous Pseudoscorpions in Texas and New Mexico1. Legs III and IV with more segments (6 beyond coxa) than legs Iand II (5 beyond coxa) (family Chthoniidae) 2All legs with same number of segments (either 5 or 6 beyond coxa) 42. Coxal spines only on coxa of leg I ApochthoniusCoxal spines not confined to leg I 33. Coxal spines only on coxa of leg II TyrannochthoniusCoxal spines on coxae of legs I and IIAphrastochthonius4. Tarsus of each leg divided into 2 parts, each leg with 6segments beyond coxa 5Tarsus of each leg undivided, each leg with 5 segments beyond coxa 95. Chelicera with inner margin of movable finger distinctly dentate 6Chelicera with inner margin of movable finger essentially smooth(family Garypidae)Archeolarca6. Abdomen with pleural membranes striate (family Syarinidae) ChitrellaAbdomen with pleural membranes granulate 77. Trichobothrium ib in middle of dorsum of chelal hand, isb onbase of fixed finger (family Bochicidae)LeucohyaTrichobothrium ib on base of fixed chelal finger along with isb(family Neobisiidae) 88. Movable finger of chelicera with spinneret in form of lowelevationMovable finger of chelicera with spinneret in form of long,usually branched galeaMicrobisiumTartarocreagris9. Femora of all legs similar, articulations between basifemora andtelofemora obsolete (family Cheiridiidae) 10Femora of legs I and II quite different from those of legs III andIV, articulations well developed (family Chemetidae) 1110. Movable finger of palpal chela with 2 trichobothria CheiridiumMovable finger of palpal chela with only 1 trichobothriumApocheiridium11. Chelicera with flagellum composed of 3 setae; hand of chelicerawith 4 setaeNeoallochernesChelicera with flagellum composed of 4 setae; hand of chelicerawith 5 setae 1212. Tarsus of leg IV with an acuminate tactile seta DinocheirusTarsus of leg IV without an acuminate tactile setaHesperochernes128
Species of Aphrastochthonius are known fromAlabama, Cuba, Central America, New Mexico, andCalifornia. Except for 2 species from Cuba andChiapas, Mexico, all are cavernicoles.Aphrastochthonius pachysetus MuchmoreAphrastochthonius pachysetus Muchmore, 1976:361, Figs. 1-3.The holotype female of this species was found inDoc Brito Cave, near White's City, Eddy County,New Mexico. No other representative is known.Genus Apochthonius ChamberlinApochthonius Chamberlin, 1929a:66; Muchmoreand Benedict, 1976:68.Apochthonius is distributed over most of theUnited States and Canada. Representatives are to befound in moist litter, both epigean and hypogean.Apochthonius magnanimus HoffApochthonius magnanimus Hoff, 1956a:4, Figs.2-4; Hoff, 1958:6.A male belonging to Apochthonius was collectedin Ogle Cave, Carlsbad Caverns National Park,Eddy County, New Mexico, 25 May 1974, by W.C.Welbourn. Though it is a little smaller than thetypes described by Hoff, it is, no doubt, arepresentative of A. magnanimus (see Welbourn,1978:31), accidental in the cave.Genus Tyrannochthonius ChamberlinTyrannochthonius Chamberlin, 1929a:74; Muchmore,1986:20.This genus has a wide distribution in the wannerparts of the world; in North America it is foundthrough the southern United States and into the Caribbeanarea and Central America (mostly unpublishedrecords). While Chamberlin and Malcolm(1960) made mention of troglobitic forms ofTyrannochthonius from Alabama, only 2 namedspecies have been reported from the United States,an epigean fonn from Florida and a troglobite fromTexas. Epigean representatives are found In moistlitter.Tyrannochthonius troglodytes MuchmoreTyrannochthonius troglodytes Muchmore, 1986:23,Fig. 11.The holotype and allotype of this species werecollected in Rock Slab Cave, on Enchanted Rock,Llano County, Texas. No other representative isknown.Tyrannochthonius texanus, new speciesFigs. 1, 2Tyrannochthonius spp.: Reddell, 1970:403.Type-data.-Holotype male (WMI078.01001)and allotype female (WM1078.01002) from ToothCave, Travis County, Texas, 14 May 1966 (J. Reddell).Paratype female from Ezell's Cave, San Marcos,Hays County, Texas, 7 September 1963 (J.Reddell); paratype male from Deep Cave, EdwardsCounty, Texas, 4 September 1965 (J. Reddell); 2paratypes (1 male, 1 female) from Wren Cave, ValVerde County, Texas, 9 April 1968 (J. Reddell andT. Mollhagen). All specimens mounted on slides(FSCA).Diagnosis.-Smaller than T. troglodytes (chelalength
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PREFACEThe present volume is the se
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TABLE OF CONTENTSHOLSINGER, JOHN R.
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the state of Coahuila in northern M
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Fig. 2.-Anesia welboumi, new specie
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\\. \ - -' ..........---~\ I// --..
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Mexiweckelia hardeni, new speciesFi
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2 sets of 1 or 2 setae each; dactyl
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AFig. 7.-Holsingerius smaragdinus,
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have as many setae on the inner pla
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Antenna 1 about 33 % length of body
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Of biogeographic interest for the h
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Bowman, T .E. 1992. Two subterranea
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A note by Scott Harden that accompa
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~~..~ ~~ I))\\ "-,'.=bFig. 3.-Speoc
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unarmed, except in pleopod 2, which
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Cokendolpher, LC., and l.R. Reddell
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lobes. The larger setae vary greatl
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the relationships of the order and
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zomids their absence is considered
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some species could be either split,
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have small pores over the surface o
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inhabits tropical deciduous forest
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huitvnolotitlensis from A. stygius
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8-10, figs. 5-7; Rowland, 1973c:136
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(in row) and one pair large posteri
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(0.34); tarsus 0.64 (0.58); total 4
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Abdomen: Tergite I with two pairs a
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setae near posterior margin. stemit
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setae, and ten ventral setae. Stemi
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1,980 m elev., 26 Dec. 1986 (T. Tre
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asitarsal-tarsal proportions: 15:4:
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and one pair setae at base of proce
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Cephalothorax: Propeltidium 1.66 mm
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Male adult unknown.Immature paratyp
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IV: trochanter 1.20 (1.10); femur 3
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Cokendolpher, 1.C. 1981. The order
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Gertsch, W.J. 1992. Distribution pa
Page 87 and 88: same families and genera, but also
Page 89 and 90: species from the United States and
Page 91 and 92: the posterior median pair short and
Page 93 and 94: great many species of North America
Page 95 and 96: Key to the Eyed Females1. Eight eye
Page 97 and 98: 39111012Figs. 1-12.-Ventral and dor
Page 99 and 100: 1419 23Figs. 13-24.-Ventral and dor
Page 101 and 102: 27 293334Figs. 25-36.-Ventral and d
Page 103 and 104: Figs. 37-48.-Ventral and retrolater
Page 105 and 106: Cicurina blanco, new speciesFigs. 7
Page 107 and 108: Type-data.-Female holotype from ins
Page 109 and 110: Description.-Female holotype: Lengt
Page 113 and 114: lengths: first femur 2 rom, fourth
Page 115 and 116: Cicurina pablo, new speciesFigs. 10
Page 119 and 120: canal in nearly vertical posItion;
Page 121 and 122: Cicurina vespera, new speciesFigs.
Page 123 and 124: procurved canal across sac; connect
Page 125 and 126: Cicurina caverna, new speciesFigs.
Page 127 and 128: with sac of similar size set in obl
Page 129 and 130: February 1964 (J. Reddell, D. McKen
Page 131 and 132: County: Diamond Cave, 16 August 196
Page 133 and 134: Roth, V.D. 1992. A new and first tr
Page 135 and 136: LITERATURE CITEDBarr, T.C. 1963. Ec
Page 137: Muchmore, W.B. 1992. Cavernicolous
Page 141 and 142: FAMILY NEOBISIIDAE CHAMBERLINGenus
Page 143 and 144: trochanter 2.5 (2.6), femur 5.55 (5
Page 145 and 146: ottom of entrance pit, Ogle Cave (O
Page 147 and 148: tactile seta on tibia and basitarsu
Page 149 and 150: Missouri may be conspecific (unpubl
Page 151 and 152: Trichobothriotaxy of chela generall
Page 153 and 154: Female (figures given first for all
Page 155 and 156: transverse furrows; eyespots not ev
Page 157 and 158: Reddell and W. Russell); I female f
Page 159 and 160: and Acuminochernes, along with the
Page 161 and 162: Chamberlin, J.C. 1946. The genera a
Page 163: (0.36); chela (without pedicel) 2.0
Page 167 and 168: although not recently studied is no
Page 169 and 170: and stylar outgrowths, present in s
Page 171 and 172: in Phalangodes (et al.) is thick an
Page 173 and 174: pairs) are found in six species: T.
Page 175 and 176: elated species IS probably best int
Page 177 and 178: legs than expected (2.6-3.2). This
Page 179 and 180: ~ 3.3 are synapomorphic); all trogl
Page 181 and 182: TAXONOMYTEXELLA Goodnight and Goodn
Page 183 and 184: 14. BK absent (Figs. 177, 180). SA
Page 185 and 186: Figs. 8-11.-Texella bijUrcata (Brig
Page 187 and 188: Description.-Total body length, 1.5
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male examined closely has fewer set
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Figs. 26-29.-Texella kokoweej, new
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apical region which loses the apica
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Figs. 38-41.-Texella shoshone, new
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Distribution.-Known only from the t
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Figs. 52-55.-Texella brevistyla, ne
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Texellajungi, new speciesFigs. 60-7
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Figs. 66-69.-Texellajungi, new spec
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than S; SA with laterobasal carina
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~81//J/'/ ;'?/ ~~.........--~~I, II
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cylindrical, retina and cornea abse
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Figs. 93-96.-Texella cokendolpheri,
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SA with well developed prong and re
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Figs. 105-108.-Texelia mulaiki Good
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and McCarty Caves, which are known
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Color orange. Body of medium rugosi
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Notes.-The type locality was errone
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Figs. 128-131.-Taella reyesi, new s
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Figs. 136-139.-Texella reyesi, new
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1989 (W. Elliott, J. Reddell, and M
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Table 3.-Continued.# locality sex S
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mesoapical; patella, 2 mesal; tibia
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Figs. 162-165.-Texella gmbbsi, new
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Figs. 166-169.-Texella diplospina,
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Figs. 174-177.-Texella renkesae, ma
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Figs. 178-18\.-Teulla spinoperca, n
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Distribution.-Known only from Fayet
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Figs. 190-193.-Texellafendi, new sp
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CLASSIFICAnONTexellabifurcata group
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Chandler, D.S. 1992. The Pselaphida
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Key to Species1. Abdominal segments
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stemite VI slightly impressed at ba
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vertexal carinae, and the laterally
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is associated with rotten woods (Ch
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small ventral carina near base, pro
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Grigarick, A.A., and R.O. Schuster.
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Cicurifla (Cicurella) holsiflgeri G
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