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spines on pleopod peduncles; 3 large basofacialspines on peduncle of uropod 1; and deep, relativelywide cleft of telson. Although some of thesesimilarities are probably more variable than others,this high number of shared characters, many ofwhich are synapomorphies, clearly indicates a muchstronger phylogenetic relationship of H.smaragdinus with H. samacos than with species ofM exiweckelia.Distribution and ecology.-This species isknown only from its type locality, where it has beencollected twice from a phreatic lake, probably exceedinga depth of 10.6 m. The lake is also inhabitedby the stygobiont cirolanid isopod Speocirolanahardeni, described by Bowman (1992, in this vol.).Etymology.-The epithet smaragdinus is fromLatin, meaning "emerald-green," and is used in referenceto the type locality which is sometimes saidto contain "emerald-green water. "FAMILY SEBIDAE WALKER, 1908Genus Seborgia BousfieldSeborgia Bousfield, 1970:164; Karaman, 1982:86-87.Relictoseborgia Karaman, 1982:91-92.Remarks on taxonomic status.-Seborgia waserected by Bousfield (1970) <strong>for</strong> a single species, S.minima, which he described from an oligohalinebrackishwater lake on Rennell Island in the BritishSolomon Islands in the South Pacific. A secondspecies, S. relicta, was described from the artesianwell in San Marcos, Texas, and assigned to this genusby Holsinger and Longley (1980).On the basis of small differences in morphologybetween these species, Karaman (1982) proposed thenew genus Relictoseborgia <strong>for</strong> S. relicta, while retainingSeborgia <strong>for</strong> S. minima. In support of hisproposal, Karaman (1982) pointed out that the Texasartesian well species differed primarily by having ashallow lateral cephalic (interantennal) lobe combinedwith an obsolete anteroventral (inferior antennal)sinus, "non-telescopically-shaped" antenna I,proportionately longer mandibular palp segment I,larger inner lobes of lower lip, unequal-sizedgnathopods, and peduncle of uropod 3 shorter thanunisegmented ramus.In my opinion most of these differences are minoror variable and all are within the accepted range ofmorphological parameters generally expected betweenspecies in the same genus (see also Holsingerand Longley, 1980; Holsinger, 1986a). However, incombination with each other and with the extremegeographic separation and significant habitat differenceof these species, I agree with Karaman thatthese differences probably support some kind of recognitionabove the species level <strong>for</strong> these taxa.However, considering the lack of strong morphologicaldifferences and the fact that only two (nowfour) species are involved, full generic status maynot be warranted. Instead, I propose dividing Seborgiainto subgenera along the lines indicated below.Subgenus Seborgia Bousfield, NEW RANKSeborgia Bousfield, 1970: 164; Karaman, 1982:86-87.Diagnosis.-Corresponding to the diagnosis <strong>for</strong>genus Seborgia given by Karaman (1982:86), exceptthat eyes may be present and the inferior antennal(anteroventral) sinus is variable in depth.Remarks.-This subgenus contains two species:S. minima Bousfield (1970), already mentionedabove; and S. schieckei Ruffo (1983) from a brackishmesopsammic habitat on the coast of South AndamanIsland in the Indian Ocean. Although S.schieckei is more closely allied morphologicallywith S. minima than S. relicta, its slightly less producedinterantennal lobe and weak inferior antennalsinus are apparently intermediate between the twoextremes noted in the other two species. To date, S.schieckei is the only eyed member of the genus.Subgenus Relictoseborgia Karaman, NEW RANKRelietoseborgia Karaman, 1982:91-92.Diagnosis.-Corresponding to the diagnosisgiven <strong>for</strong> genus Relictoseborgia by Karaman (1982).Remarks.-This subgenus contains two species:S. relicta (Holsinger, in Holsinger and Longley,1980) from the artesian well in San Marcos, HaysCo., Texas and a new locality from Medina Co.,Texas, noted below; and S. hershleri, new species,from Val Verde Co., Texas, described below.Seborgin (Relictoseborgin) hersh/eri, new speciesFig. 10Material examined.-TEXAS: Val VerdeCounty: unnamed spring on east side of DevilsRiver, ca. 32 km N of Del Rio, female holotype and3 female paratypes, R. Hershler, 8 Sept. 1986;16

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