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searchable PDF - Association for Mexican Cave Studies

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the relationships of the order and then those <strong>for</strong>members of the family. The character numbers inthe following discussion are the same as those appearingin Tables 1-2 and Fig. 2.Some authors consider the Schizomida to be asuborder of Uropygi, whereas others consider it anorder. We, like some previous authors, prefer torecognize the Schizomida and Uropygi as separateorders within the Camarostomata. Most authorscombining the two orders emphasize the existence ofplesiomorphies or characters of undetermined polarity.Some of the differences between Schizomida andUropygi are as great as the differences among theUropygi, Amblypygi, and Araneae. Shear et al.(1987) suggested that the pygidial exocrine glandwas an autapomorph <strong>for</strong> the Uropygi. This is not thecase, as such glands are also present in the Schizomida(Brignoli, 1973).Synapomorphies <strong>for</strong> the Uropygi are: pedipalpschelate, coxal gland orifices associated with leg IIIabsent, lateral tergocoxal muscle inserted on thepleural membrane adjacent to the coxa, presence ofommatoids on dorsal surface of anal somite, flagellumwith clear regions ventrally which are sensitiveto light, presence of five posterior abdominalneuromeres, and modification of cheliceral cleaningorgan.The Schizomida can be separated from theUropygi as well as other Arachnidea by the followingsynapomorphies: prosomal heart segments absent(character 0); prosoma divided into pro-,meso-, and metapeltidia (character 1); modificationof the anterior portion of the prosoma into a pointedprocess (character 2); presence of setae on and basalto the anterior process of the propeltidium (character3); median eyes absent (character 4); presence ofpairs of dorsal setae on the propeltidium (character5); stigmata lacking on segment IX (character 6);with six pairs of intestinal diverticula in the abdomen(character 7); with eight anterior abdominalneuromeres (character 8); male flagellum not dividedinto numerous segments (character 13); propeltidiumand tergites with distinct setal pattern(character 9); tergites I-II with pairs of microsetae(character 10); anterior distal tip of coxa II prolongedas a sharp process (character 11); ability tojump, femur IV and its associated muscles beingenlarged (character 12). Three additional charactersare known which appear to be synapomorphies <strong>for</strong>the Schizomida, but because they have only beendetermined (examined) in a single species of Hubbardiinaewe do not code them <strong>for</strong> the data matrix:absence of a posteriorly directed dorsal endosternitesuspensor muscle; absence of anterior transpatellarmuscle; and absence of the posterior patellotibialmuscle. Additional characters are known (Millot,1942) <strong>for</strong> which we have not determined the polarities.Specifically, uropygids have four pairs of prosomaticdiverticula, schizomids have one palr;Table 2.-Character states in selected members of thesuperorder Camarostomata (0 codes plesiomorphic condition;1,2,3 codes apomorphic conditions; ? codes unknown conditionor condition not relevant to region of cladogram being resolved).1, Uropygi; 2, Protoschizomus pachypalpus; 3, P. rowlandi; 4,P. occidentalis; 5, P. sprousei; 6, P. genschi (male unknown);7, P. purificacion (male unknown); 8, P. treacyae (maleunknown); 9, Agastoschizomus lucifer; 10, A. huitzmolotitlensis(female unknown); 11, A. srygius (male unknown); 12, A. patei;13, Megaschizominae; 14, Hubbardiinae.TaxaI 2 3 4 5 6 7 8 9 10 11 12 13 14Character0.0 1 I I 1 1 1 1 I I 1 1I. 0 1 I I I 1 I I I I I I2.0 I I I 1 I 1 1 I I I I3.0 I I 1 1 1 1 I 1 1 1 14.0 I I 1 1 I I 1 1 1 1 15.0 1 1 I I 1 1 1 1 I 1 16.0 1 1 1 1 1 1 1 I 1 I 17. 0 I I 1 1 I 1 I I I I I8.0 1 1 1 1 1 I 1 1 1 1 19.0 1 I 1 1 1 I I I 1 1 I10.0 1 1 I 1 I 1 1 1 I 1 111. 0 I I I I 1 1 I I I I I12.0 1 I 1 I 1 1 1 1 I 1 I13.0 1 I 1 I ? ? ? I I ? I I 114.0 0 0 0 0 0 0 0 0 0 0 0 1 2IS.? 0 0 0 0 0 0 0 0 0 0 0 1 116.0 0 0 0 0 0 0 0 0 0 0 0 1 117.? 0 0 0 0 0 0 0 1 1 1 1 2 0,318.0 1 1 1 1 1 1 1 1 1 1 1 0 019.? 1 I 1 0 0 0 0 0 0 0 0 0 0,220.0 1 I II? ? ? 1 ? ? 1 0 021.? 1 1 1 0 0 0 0 0 0 0 0 ? ?22.? 0 0 0 I 0 1 0 0 0 0 0 ? ?23.? 0 0 0 0 0 0 0 0 0 0 0 1 224.? 0 0 0 0 0 0 0 I I I I 0 025.0 I I I 1 1 I I 0 0 0 0 2 026.? I 1 1 0 0 0 0 0 0 0 0 ? ?27.? 0 0 0 0 ? ? ? II? I 0 028.? I I I 1 I 1 1 0 0 0 0 0 029.0 0 0 0 0 0 0 0 0 0 0 0 1 130.? 0 0 ? 0 0 0 0 0 ? 0 2 0 I31.? I I ? I 1 I II? 1 I 0 0,232.? I I ? I ?? 1 1 0 133.? 0 0 ? 0 0 0 0 ? 2 0 0 034.? 0 0 0 0 O? 1 I ? 1 0 035.? I I 0 0 0 0 0 0 0 0 0 ? ?36.? 0 0 ? I 0 I 0 0 ? 0 0 0 037.? 1 I 1 I ? ? ? 0 0 ? 0 0 0,238.? 1 I 1 I ? ? ? II? 1 0 039.? 0 0 0 0 ? ? ? II? 1 0 0,240.? I I 0 0 ? ? ? 0 0 ? 0 ? ?41.? 0 0 0 1 ? ? ? 0 2 ? 0 ? ?42.? 0 0 0 0 0 0 0 0 ? 0 0 I 243.? 1 1 1 0 0 0 0 0 ? 3 2 ? ?35

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