and tibia 5.4 times as long as deep. Tarsus of leg IVwith a short, acuminate tactile seta 3/4 length ofsegment from proximal end.Measurements (mm).-Body length 2.75. Carapacelength 0.805. Chelicera length 0.275. Palpaltrochanter 0.435 by 0.225; femur 0.715 by 0.235;tibia 0.62 by 0.26; chela (without pedicel) 1.18 by0.375; hand (without pedicel) 0.58 by 0.35; pedicel0.13 long; movable finger length 0.62. Leg IV: entirefemur 0.65 by 0.155; tibia 0.54 by 0.10; tarsus0.43 by 0.08.Etymology.-The species is named incertus becauseof its uncertain taxonomic position.Remarks.-This species is tentatively placed inNeoallochernes on the basis of these characters: 3setae in the cheliceral flagellum; only 4 setae oncheliceral hand, sbs lacking; location of "tactileseta" on tarsus of leg IV at distal 3/4; and possessionof a vestigial venom duct in the fixed chelalfinger. However, it differs from others in that genusin several important characters and may well belongto an hitherto undescribed genus. Some differencesfrom other known members of Neoallochernes arethe following: on the cheliceral hand, seta bs isacuminate rather than denticulate and es is shorterthan bs rather than longer; on the tarsus of leg IVthe "tactile seta" is acuminate rather than denticulate;on the movable chelal finger, the distance betweentrichobothria t and st is twice as great as thedistance between band sb rather than the 2 distancesessentially equal, and the nodus ramosus of thevenom dust is midway between t and st rather thanproximad of st; on the fixed chelal finger, est is distadof the middle rather than proximad, and thegroup eb, esb, ib and isb lies mostly distal to thelevel of the last marginal tooth rather than proximalto the last tooth; and eyespots are apparently absentrather than present. These differences probably indicatethat the new species represents a new genus, butI prefer not to name it now on the basis of a single,damaged male specimen. The differences should beconfirmed on additional material and the nature ofthe genitalia, especially of the female, should beelucidated. At hand are a few specimens from CentralAmerica which appear to be congeneric and mayallow a proper characterization of the taxon.Genus Hesperochernes ChamberlinHesperochernes Chamberlin, 1924b:89; Muchmore,1974:27.Representatives of the genus Hesperochernes arewidely distributed through the temperate andtropical parts of North and Central America. Threespecies have been reported previously from NewMexico, but only one from Texas (Hoff, 1958).Many of the <strong>for</strong>ms are found in close associationwith small mammals, especially packrats, andseveral are recorded from caves, usually on batguano.Hesperochernes occidentalis (Hoff and Bolsterli)Pseudozaona occidentalis Hoff and Bolsterli,1956: 170, figs. 1-3; Hoff, 1958:24.Hesperochernes occidentalis (Hoff and Bolsterli):Muchmore, 1974:30.This species was first described from caves inWashington County, Arkansas. It has since beenfound in many other caves in the Ozark Region ofArkansas, Missouri and Oklahoma (unpublished).There are 2 collections of H. occidentalis fromTexas, both from Edwards County: 4 specimens (1male, 2 females, 1 tritonymph) from Wyatt <strong>Cave</strong>, 2miles north of Wheat <strong>Cave</strong>, 21 September 1963 (J.Reddell and D. McKenzie); 3 specimens (1 male, 1female, 1 tritonymph from nest of Petrochelidon fulva(Vieillot) in Dunbar <strong>Cave</strong>, 22 July 1976 (R.Martin). The species is highly variable in size andproportions and these individuals fit easily into theknown ranges of measurements.Hesperochernes riograndensis Hoff and ClawsonHesperochernes riograndensis Hoff and Clawson,1952:19, figs. 11-12; Hoff, 1958:23.A number of specimens collected from caves inwestern Texas and eastern New Mexico appear tobelong to this species, which has been known previouslyonly from "food storage of a kangaroo rat(Dipodomys) " in Socorro County, New Mexico(Hoff and Clawson, 1952:23). Most of them arelarger than the types described by Hoff and Clawson,but they have the same general shape and proportions.On the other hand, some of them havesome characters similar to those of H. canadensisHoff and H. utahensis Hoff and Clawson, both ofwhich have been reported in the southern RockyMountain region. Until a comprehensive revision ofthe western species of Hesperochernes is completed,it is impossible to be certain of the identities of mostindividuals.Present collections include: 1 female from"below dome 250 feet from entrance" in Blackstone<strong>Cave</strong>, Terrell County, Texas, 4 June 1963 (J.146
Reddell and W. Russell); I female from MurphyWells <strong>Cave</strong>, Irion County, Texas, 24 February 1974(R. Ballinger); 3 males from dark zone of Wind<strong>Cave</strong> (upper level), Eddy County, New Mexico, 31December 1973 (W.C. Welbourn); 1 female fromHelen's <strong>Cave</strong>, Carlsbad <strong>Cave</strong>rns National Park,Eddy County, New Mexico, 31 August 1974 (W.e.Welbourn); 1 female near bat guano in Endless<strong>Cave</strong>, Eddy County, New Mexico, 17 February1975 (W.C. Welbourn); 3 males, 1 female fromThree Fingers <strong>Cave</strong>, Lincoln National Forest, EddyCounty, New Mexico, 28 May 1978 (W.C. Welbourn);1 female in "twilight zone near porcupinelair," Serpentine Root <strong>Cave</strong>, NW Tinnie, LincolnCounty, New Mexico, 27 July 1973 (W.C. Welbourn).All are mounted on slides (FSCA).Hesperochernes molestus HoffHesperochernes molestus Hoff, 1956c:33, figs.12-15; Hoff, 1958:24; Hoff, 1959:32.?Hesperochernes sp.: Reddell, 1970:403.This species has been known previously onlyfrom central New Mexico; found in close associationwith rodents, in nests of Neotoma sp., Dipodomyssp., and Perognathus flavus Baird, and onthe hair of Onychomys leucogaster (Wied-Neuwied)(Hoff, 1956c:38).At hand is one collection of 26 specimens (21males, 4 females, 1 deutonymph) which appear tobelong to this species. They were taken from "batguano in the Bat Room," Four Mile <strong>Cave</strong>, 4 milesN Del Rio, Val Verde County, Texas, 24 April1966 (1. Reddell and E. Alexander); 10 adults (6males, 4 females) mounted on slides (FSCA). Theytend to be a little larger on average than the typesreported by Hoff from New Mexico, but in generalshape and proportions they are much the same.A single conspecific female was found "attachedto the right rear leg of a cranefly" at College Station,Brazos County, Texas, II March 1985 (S.W.Taber). This sort of phoretic association may assist,at least in part, in the spread of the species fromplace to place.Hesperochernes unicolor (Banks)Chelanops unicolor Banks, 1908:39.Hesperochernes unicolor (Banks): Hoff, 1947:5II;Hoff and Clawson, 1952:23.A single female referable to this species wasfound in Ezell's <strong>Cave</strong>, Hays County, Texas, 15August 1978 (J.e. Davis); mounted on slide(FSCA). It is a little larger than 3 femalespreviously reported by Hoff (1947) and Hoff andClawson (1952), but otherwise fits the description ofthe species very well. It was obviously accidental inthe cave, not at all modified <strong>for</strong> cave life.Genus Dinocheirus ChamberlinDinocheirus Chamberlin, 1929b: 171; Muchmore,1974:31.Dinocheirus, as presently (imperfectly) understood,is widely distributed in Europe and North andCentral America. Five species have been reportedpreviously from New Mexico and 2 from Texas(Hoff, 1958). Representatives are usually found inrich <strong>for</strong>est litter or in the nests of small mammals;one species has been reported from bat guano inCarlsbad <strong>Cave</strong>rns.Dinocheirus astutus HoffDinocheirus astutus Hoff, 1956c:44, figs. 18-20;Hoff, 1958:28; Hoff, 1959:27, 33; Barr andReddell, 1967:259.Dinocheirus astutus was described by Hoff(1956c) from specimens taken mainly from nests ofNeotoma sp. in west-central New Mexico. It wassubsequently reported (del. Hoff) from the Bat<strong>Cave</strong>, Carlsbad <strong>Cave</strong>rns, in southeastern New Mexico(Barr and Reddell, 1967). More recent collectionshave provided many more specimens fromCarlsbad <strong>Cave</strong>rns and a single female from Ellis<strong>Cave</strong> in west-central New Mexico.Nine males and 10 females from the recent collectionshave been mounted and studied. In mostparticulars they agree closely with Hoff's descriptions.However, both males and females tend to be alittle larger and have slightly more slender appendagesthan Hoff's. These differences may be an artifactof different techniques of measurement or mayindicate some small adaptations of the cavernicolous<strong>for</strong>ms.New records.-NEW MEXICO: Eddy County:Carlsbad <strong>Cave</strong>rns National Park, Bat <strong>Cave</strong>, berlesesample of guano of Tadarida brasiliensis, 25-27November 1967 (T. Rossen), about 150 specimens,all stages; berlese samples, 14 February 1974, 27June 1974, 28 November 1974, 26-27 April 1975(W.C. WeIbourn), about 150 specimens, all stages.147
- Page 7 and 8:
PREFACEThe present volume is the se
- Page 9:
TABLE OF CONTENTSHOLSINGER, JOHN R.
- Page 12 and 13:
the state of Coahuila in northern M
- Page 14 and 15:
Fig. 2.-Anesia welboumi, new specie
- Page 16 and 17:
\\. \ - -' ..........---~\ I// --..
- Page 18 and 19:
Mexiweckelia hardeni, new speciesFi
- Page 20:
2 sets of 1 or 2 setae each; dactyl
- Page 23 and 24:
AFig. 7.-Holsingerius smaragdinus,
- Page 25 and 26:
have as many setae on the inner pla
- Page 28:
Antenna 1 about 33 % length of body
- Page 31 and 32:
Of biogeographic interest for the h
- Page 33 and 34:
Bowman, T .E. 1992. Two subterranea
- Page 35 and 36:
A note by Scott Harden that accompa
- Page 37 and 38:
~~..~ ~~ I))\\ "-,'.=bFig. 3.-Speoc
- Page 39 and 40:
unarmed, except in pleopod 2, which
- Page 41 and 42:
Cokendolpher, LC., and l.R. Reddell
- Page 43 and 44:
lobes. The larger setae vary greatl
- Page 45 and 46:
the relationships of the order and
- Page 47 and 48:
zomids their absence is considered
- Page 49 and 50:
some species could be either split,
- Page 51 and 52:
have small pores over the surface o
- Page 53 and 54:
inhabits tropical deciduous forest
- Page 55 and 56:
huitvnolotitlensis from A. stygius
- Page 57 and 58:
8-10, figs. 5-7; Rowland, 1973c:136
- Page 59 and 60:
(in row) and one pair large posteri
- Page 61 and 62:
(0.34); tarsus 0.64 (0.58); total 4
- Page 63 and 64:
Abdomen: Tergite I with two pairs a
- Page 65 and 66:
setae near posterior margin. stemit
- Page 67:
setae, and ten ventral setae. Stemi
- Page 71 and 72:
1,980 m elev., 26 Dec. 1986 (T. Tre
- Page 73 and 74:
asitarsal-tarsal proportions: 15:4:
- Page 75 and 76:
and one pair setae at base of proce
- Page 77 and 78:
Cephalothorax: Propeltidium 1.66 mm
- Page 79 and 80:
Male adult unknown.Immature paratyp
- Page 81 and 82:
IV: trochanter 1.20 (1.10); femur 3
- Page 83 and 84:
Cokendolpher, 1.C. 1981. The order
- Page 85 and 86:
Gertsch, W.J. 1992. Distribution pa
- Page 87 and 88:
same families and genera, but also
- Page 89 and 90:
species from the United States and
- Page 91 and 92:
the posterior median pair short and
- Page 93 and 94:
great many species of North America
- Page 95 and 96:
Key to the Eyed Females1. Eight eye
- Page 97 and 98:
39111012Figs. 1-12.-Ventral and dor
- Page 99 and 100:
1419 23Figs. 13-24.-Ventral and dor
- Page 101 and 102:
27 293334Figs. 25-36.-Ventral and d
- Page 103 and 104:
Figs. 37-48.-Ventral and retrolater
- Page 105 and 106: Cicurina blanco, new speciesFigs. 7
- Page 107 and 108: Type-data.-Female holotype from ins
- Page 109 and 110: Description.-Female holotype: Lengt
- Page 111 and 112: Description.-Female holotype: Lengt
- Page 113 and 114: lengths: first femur 2 rom, fourth
- Page 115 and 116: Cicurina pablo, new speciesFigs. 10
- Page 117 and 118: Description.-Female holotype: Lengt
- Page 119 and 120: canal in nearly vertical posItion;
- Page 121 and 122: Cicurina vespera, new speciesFigs.
- Page 123 and 124: procurved canal across sac; connect
- Page 125 and 126: Cicurina caverna, new speciesFigs.
- Page 127 and 128: with sac of similar size set in obl
- Page 129 and 130: February 1964 (J. Reddell, D. McKen
- Page 131 and 132: County: Diamond Cave, 16 August 196
- Page 133 and 134: Roth, V.D. 1992. A new and first tr
- Page 135 and 136: LITERATURE CITEDBarr, T.C. 1963. Ec
- Page 137 and 138: Muchmore, W.B. 1992. Cavernicolous
- Page 139 and 140: Species of Aphrastochthonius are kn
- Page 141 and 142: FAMILY NEOBISIIDAE CHAMBERLINGenus
- Page 143 and 144: trochanter 2.5 (2.6), femur 5.55 (5
- Page 145 and 146: ottom of entrance pit, Ogle Cave (O
- Page 147 and 148: tactile seta on tibia and basitarsu
- Page 149 and 150: Missouri may be conspecific (unpubl
- Page 151 and 152: Trichobothriotaxy of chela generall
- Page 153 and 154: Female (figures given first for all
- Page 155: transverse furrows; eyespots not ev
- Page 159 and 160: and Acuminochernes, along with the
- Page 161 and 162: Chamberlin, J.C. 1946. The genera a
- Page 163: (0.36); chela (without pedicel) 2.0
- Page 167 and 168: although not recently studied is no
- Page 169 and 170: and stylar outgrowths, present in s
- Page 171 and 172: in Phalangodes (et al.) is thick an
- Page 173 and 174: pairs) are found in six species: T.
- Page 175 and 176: elated species IS probably best int
- Page 177 and 178: legs than expected (2.6-3.2). This
- Page 179 and 180: ~ 3.3 are synapomorphic); all trogl
- Page 181 and 182: TAXONOMYTEXELLA Goodnight and Goodn
- Page 183 and 184: 14. BK absent (Figs. 177, 180). SA
- Page 185 and 186: Figs. 8-11.-Texella bijUrcata (Brig
- Page 187 and 188: Description.-Total body length, 1.5
- Page 189 and 190: male examined closely has fewer set
- Page 191 and 192: Figs. 26-29.-Texella kokoweej, new
- Page 193 and 194: apical region which loses the apica
- Page 195 and 196: Figs. 38-41.-Texella shoshone, new
- Page 197 and 198: Distribution.-Known only from the t
- Page 199 and 200: Figs. 52-55.-Texella brevistyla, ne
- Page 201 and 202: Texellajungi, new speciesFigs. 60-7
- Page 203 and 204: Figs. 66-69.-Texellajungi, new spec
- Page 205 and 206: than S; SA with laterobasal carina
- Page 207 and 208:
~81//J/'/ ;'?/ ~~.........--~~I, II
- Page 209 and 210:
cylindrical, retina and cornea abse
- Page 211 and 212:
Figs. 93-96.-Texella cokendolpheri,
- Page 213 and 214:
SA with well developed prong and re
- Page 215 and 216:
Figs. 105-108.-Texelia mulaiki Good
- Page 217 and 218:
and McCarty Caves, which are known
- Page 219 and 220:
Color orange. Body of medium rugosi
- Page 221 and 222:
Notes.-The type locality was errone
- Page 223 and 224:
Figs. 128-131.-Taella reyesi, new s
- Page 225 and 226:
Figs. 136-139.-Texella reyesi, new
- Page 227 and 228:
Figs. 144-147.-Texella reyesi, new
- Page 229 and 230:
Figs. 152-155.-Texella reyesi, new
- Page 231 and 232:
1989 (W. Elliott, J. Reddell, and M
- Page 233 and 234:
Table 3.-Continued.# locality sex S
- Page 235 and 236:
mesoapical; patella, 2 mesal; tibia
- Page 237 and 238:
Figs. 162-165.-Texella gmbbsi, new
- Page 239 and 240:
Figs. 166-169.-Texella diplospina,
- Page 241 and 242:
Figs. 174-177.-Texella renkesae, ma
- Page 243 and 244:
Figs. 178-18\.-Teulla spinoperca, n
- Page 245 and 246:
Distribution.-Known only from Fayet
- Page 247 and 248:
Figs. 190-193.-Texellafendi, new sp
- Page 249 and 250:
CLASSIFICAnONTexellabifurcata group
- Page 251 and 252:
Chandler, D.S. 1992. The Pselaphida
- Page 253 and 254:
Key to Species1. Abdominal segments
- Page 255 and 256:
stemite VI slightly impressed at ba
- Page 257 and 258:
vertexal carinae, and the laterally
- Page 259 and 260:
is associated with rotten woods (Ch
- Page 261 and 262:
small ventral carina near base, pro
- Page 263:
Grigarick, A.A., and R.O. Schuster.
- Page 266 and 267:
Cicurifla (Cicurella) holsiflgeri G