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searchable PDF - Association for Mexican Cave Studies

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Cokendolpher, LC., and l.R. Reddell. 1992. Revision of the Protoschizomidae (Arachnida: Schizomida) with notes on the phylogenyof the order. Texas Mem. Mus., Speleol. Monogr., 3:31-74REVISION OF THE PROTOSCHIZOMIDAE (ARACHNIDA: SCHIZOMIDA)WITH NOTES ON THE PHYLOGENY OF THE ORDERJames C. Cokendolpher2007 29th St.Lubbock, TX 79411andJames R. ReddellTexas Memorial MuseumThe University of Texas at Austin2400 TrinityAustin, TX 78705ABSTRACTThe predominantly <strong>Mexican</strong> family Protoschizomidae isrevised to include 11 species in two genera. ProroschizomusRowland is recognized <strong>for</strong> seven species, five of which areplaced in two species groups: the pachypalpus group includes P.pachypalpus (Rowland) (Tamaulipas), P. rowlandi n. sp. (SanLuis Potosi), and P. occidenralis Rowland (Colima); the sprouseigroup includes P. sprousei n. sp. and P. purificacion n. sp. (bothfrom Tamaulipas). Two species known only from females, P.treacyae n. sp. and P. gertschi n. sp. (both from Tamaulipas),are not placed in species groups. Agastoschizomus Rowlandincludes A. lucifer Rowland (San Luis Potosi), A. huitzmolotitlensisRowland (San Luis Potosi), A. stygius n. sp. (Hidalgo),and A. patei (Tamaulipas). Several undetermined immaturespecimens, including the first species of the family from Texasand the U.S.A., are also recorded. The female genitalia aredescribed <strong>for</strong> the first time <strong>for</strong> members of the Protoschizomidae.A new system <strong>for</strong> naming articles in the female flagellum and arevised method <strong>for</strong> numbering flagellar setae are provided. Thephylogeny of the order Schizomida is discussed, and a cladogramof the families and protoschizomid genera and species isprovided. The available data on habitats and cave life arerecorded. Agasroschizomus patei may be paedomorphic, the first<strong>for</strong> the order.INTRODUCTIONThe family Protoschizomidae is a small distinctivegroup of the order Schizomida known only fromsouthwestern Texas (U.S.A.) and Mexico. The firstmember of the family Protoschizomidae was describedby Rowland (1971) as Agastoschizomus luciferand placed in the Hubbardiidae (=Schizomidae).The subfamily Megaschizominae was named byRowland (1973b) to include Agastoschizomus andthe African genus Megaschizomus Lawrence. Rowland(1975b) later recognized the distinctness of theAmerican <strong>for</strong>ms and proposed the family Protoschizomidaeto include A. lucifer, A. huitzmolotitlensisRowland, Protoschizomus pachypalpus Rowland,and P. occidentalis Rowland. Megaschizomus wasretained in the monotypic subfamily Megaschizominaein the Hubbardiidae (=Schizomidae).In the present paper we redescribe the knownspecies, describe five new species of Protoschizomus31

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