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9.(6) Antennomeres III-VIII twice as long as wide; sides of head ineye region gently curved (Fig. 9)5.Batrisodes texanus n. sp.At least antennomere VIII less than twice as long as wide; lateralmargins of head slightly angulate and bearing granules whereeyes should be (Figs. 7, 13) 1010.(9) Head lacking lateral and medial vertexal carinae (Fig. 13),vertex smoothHead with at least lateral carinae extending anteriorly from base(Figs. 5, 7), vertex in apical half distinctly punctate or rugulose11.(10) Apical half of vertex roughened (females) or coarsely punctate(males), with faint lateral vertexal carinae, lacking medianvertexal carina; males with anterior margin of vertex broadlytruncate, first antennomere not modified (Fig. 5)Apical half of vertex smooth (females) to transversely rugulose(males), with median and lateral vertexal carinae sharplydefined; males with anterior margin of vertex broadlybisinuate, first antennomere angulate ventrally (Fig. 7)6. Batrisodes venyivi n.sp.ll3. Batrisodes grubbsi n.sp.4. Batrisodes reyesi n. sp.these have been collected in sawdust, under pinebark, and under stones. This species has not beenpreviously noted as a troglophile, and it is interestingthat in surface habitats this species is alwayscollected in quite short series. The longest series Ihave seen is from Ireland's Cave.James R. Reddell (pers. comm.) has supplied thefollowing collection information. "It has been foundbelow rocks in moist sinkhole-type caves in associationwith troglobitic millipedes and other species,but also occurs in total darkness. Most specimenshave been found on the underside of rocks lightlyburied in silt. A specimen from Beck Ranch Cavewas found crawling on moist flowstone more than200 meters from the cave entrance. "Park (1960) described a single female fromSchneider Cave as a new species, schneiderensis.Several series from Texas caves contain both sexes,and it is obvious that the type of schneiderensis is atypical female of uncicornis. Batrisodes uncicornishas been placed close to riparius (Say) based on thevery similar male antennal characters (Park, 1947).The modifications of the legs are also similar, as isthe general structure of the head. The punctation ofthe head is coarser in uncicornis, and the apical angieof the vertex is only slightly depressed, while inriparius the punctation is not coarse on the pronotumand basal portion of the vertex, and the apicalportion of the vertex is strongly deflexed betweenthe antennal bases. Females of uncicornis may beconfused with those of globosus (LeConte), but maybe separated by the head being more completely andcoarsely punctate.2. Batrisodes (Excavodes) clypeonotus (Brendel)(Figs. 3-4)Batrisus clypeonotus Brendel, 1893:280, pI. IV.Type locality: Ponchatoula, Louisiana. Maleholotype (ANSP).Batrisodes tridens Casey, 1908:263. Type locality:St. Louis, Missouri. Male holotype (USNM).NEW SYNONYMY.Description.-Length 2.28-2.68. Males withhead transversely excavate just anterior to antennalbases, frons with acute medial and lateral tubercles,medial tubercle with short posterolaterally directedsetae, lateral tubercles glabrous, vertex sharply angledventrally between antennal bases, this area setwith short obvious setae arising from small distinctpunctures, vertexal apex with small median trilobedprojection bearing two small tufts of setae, basalportion of vertex smoother, sparsely setate withsetae arising from isolated granules, vertexal foveaenude, lateral vertexal carinae weak and extending toantennal bases, median carina weak and extending tomiddle of vertex, eyes with 12 (Texas) to 40(Missouri) facets grouped in crescentic pattern; firstantennomere (scape) enlarged, impressed at center ofanterior face, densely and minutely punctate in impression,antennomere X globose, slightly widerthan last antennomere and lacking a ventral fovea.Pronotum with median longitudinal sulcus extendingto near apex, disc sparsely punctate; elytrawith three basal foveae and subhumeral fovea;244
stemite VI slightly impressed at base with discrounded; second mesotarsomeres straight.Females with frons and anterior half of vertexminutely rugose, small eyes with 10-16 facets, lackingmodifications of head and antennae, stemite VIbroadly rounded.Male from Powell's Cave, Texas: antennae 1.08,metafemora 0.80, metatibiae 0.77, metatarsi 0.45.Specimen examined.-TEXAS: Menard County:Powell's Cave, IV-II-1989, E. Shoud, third crevice(TMMC).Comments.-This species has been poorlyknown since its description, and during preparationof this paper was found to be the senior synonym oftridens Casey. Park (1956:84) examined a specimenhe placed as this species from northern Alabama,and found a minute fovea of antennomere X. A slidepreparation of a specimen from Oklahoma did notreveal any trace of a fovea, and a fovea is not obviousin any of the specimens from several localitiesmounted on points. Park (1956) believed thatclypeonotus represented an undescribed subgenus,but it appears typical of Excavodes to me. Thisspecies is now known from Missouri to Alabamaand Texas. The largest series have been generatedby collecting specimens with their ant host, Camponotusamericanus Mayr. Whether this species isan obligate myrmecophile is not known, but thereduced number of eye facets indicates that clypeonotushas adapted to a subterranean life.3. Batrisodes (Excavodes) grubbsi n. sp.(Figs. 5-6)Description.-Length 2.32-2.48. Male head withtransverse excavation anterior to antennal bases,clypeus with erect blunt tubercle, tubercle denselysetate along postero-Iateral margin, vertex broadlytruncate in dorsal view, in anterior view vertex projectingventrally to medial point that is slightly angledanteriorly near apex, vertexal apex densely setateinto transverse excavation, vertex coarselypunctate, smoothly roughened in area betweenvertexal foveae, median carina faint, lateral carinaelow but distinct and extending to just anterior toeyes, eyes consisting of approximately 8 granulesFigs. 1-7.-Batrisodes spp.: I, B. uncicomis, habitus of male and anterior view, right antenna; 2, B. uncicomis, dorsal and rightlateral view, aedeagus; 3, B. clypeonotus, head and anterior view, right antenna of male; 4, B. clypeonotus, dorsal and right lateralview, aedeagus; 5, B. grubbsi, head and anterior view, right antenna of male; 6, B. grubbsi, dorsal and right lateral view, aedeagus; 7,B. reyesi, head and anterior view, right antenna of male. Scale line equals O. J. Apical three segments of male antennae presented inventral view.245
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PREFACEThe present volume is the se
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TABLE OF CONTENTSHOLSINGER, JOHN R.
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the state of Coahuila in northern M
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Fig. 2.-Anesia welboumi, new specie
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\\. \ - -' ..........---~\ I// --..
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Mexiweckelia hardeni, new speciesFi
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2 sets of 1 or 2 setae each; dactyl
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AFig. 7.-Holsingerius smaragdinus,
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have as many setae on the inner pla
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Antenna 1 about 33 % length of body
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Of biogeographic interest for the h
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Bowman, T .E. 1992. Two subterranea
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A note by Scott Harden that accompa
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~~..~ ~~ I))\\ "-,'.=bFig. 3.-Speoc
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unarmed, except in pleopod 2, which
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Cokendolpher, LC., and l.R. Reddell
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lobes. The larger setae vary greatl
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the relationships of the order and
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zomids their absence is considered
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some species could be either split,
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have small pores over the surface o
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inhabits tropical deciduous forest
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huitvnolotitlensis from A. stygius
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8-10, figs. 5-7; Rowland, 1973c:136
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(in row) and one pair large posteri
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(0.34); tarsus 0.64 (0.58); total 4
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Abdomen: Tergite I with two pairs a
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setae near posterior margin. stemit
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setae, and ten ventral setae. Stemi
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1,980 m elev., 26 Dec. 1986 (T. Tre
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asitarsal-tarsal proportions: 15:4:
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and one pair setae at base of proce
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Cephalothorax: Propeltidium 1.66 mm
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Male adult unknown.Immature paratyp
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IV: trochanter 1.20 (1.10); femur 3
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Cokendolpher, 1.C. 1981. The order
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Gertsch, W.J. 1992. Distribution pa
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same families and genera, but also
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species from the United States and
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the posterior median pair short and
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great many species of North America
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Key to the Eyed Females1. Eight eye
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39111012Figs. 1-12.-Ventral and dor
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1419 23Figs. 13-24.-Ventral and dor
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27 293334Figs. 25-36.-Ventral and d
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Figs. 37-48.-Ventral and retrolater
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Cicurina blanco, new speciesFigs. 7
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Type-data.-Female holotype from ins
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Description.-Female holotype: Lengt
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lengths: first femur 2 rom, fourth
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Cicurina pablo, new speciesFigs. 10
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canal in nearly vertical posItion;
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Cicurina vespera, new speciesFigs.
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procurved canal across sac; connect
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Cicurina caverna, new speciesFigs.
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with sac of similar size set in obl
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February 1964 (J. Reddell, D. McKen
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County: Diamond Cave, 16 August 196
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Roth, V.D. 1992. A new and first tr
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LITERATURE CITEDBarr, T.C. 1963. Ec
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Muchmore, W.B. 1992. Cavernicolous
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Species of Aphrastochthonius are kn
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FAMILY NEOBISIIDAE CHAMBERLINGenus
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trochanter 2.5 (2.6), femur 5.55 (5
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ottom of entrance pit, Ogle Cave (O
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tactile seta on tibia and basitarsu
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Missouri may be conspecific (unpubl
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Trichobothriotaxy of chela generall
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Female (figures given first for all
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transverse furrows; eyespots not ev
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Reddell and W. Russell); I female f
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and Acuminochernes, along with the
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Chamberlin, J.C. 1946. The genera a
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(0.36); chela (without pedicel) 2.0
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although not recently studied is no
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and stylar outgrowths, present in s
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in Phalangodes (et al.) is thick an
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pairs) are found in six species: T.
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elated species IS probably best int
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legs than expected (2.6-3.2). This
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~ 3.3 are synapomorphic); all trogl
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TAXONOMYTEXELLA Goodnight and Goodn
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14. BK absent (Figs. 177, 180). SA
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Figs. 8-11.-Texella bijUrcata (Brig
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Description.-Total body length, 1.5
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male examined closely has fewer set
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Figs. 26-29.-Texella kokoweej, new
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apical region which loses the apica
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Figs. 38-41.-Texella shoshone, new
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Distribution.-Known only from the t
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Figs. 52-55.-Texella brevistyla, ne
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Texellajungi, new speciesFigs. 60-7
Page 203 and 204: Figs. 66-69.-Texellajungi, new spec
Page 205 and 206: than S; SA with laterobasal carina
Page 207 and 208: ~81//J/'/ ;'?/ ~~.........--~~I, II
Page 209 and 210: cylindrical, retina and cornea abse
Page 211 and 212: Figs. 93-96.-Texella cokendolpheri,
Page 213 and 214: SA with well developed prong and re
Page 215 and 216: Figs. 105-108.-Texelia mulaiki Good
Page 217 and 218: and McCarty Caves, which are known
Page 219 and 220: Color orange. Body of medium rugosi
Page 221 and 222: Notes.-The type locality was errone
Page 223 and 224: Figs. 128-131.-Taella reyesi, new s
Page 225 and 226: Figs. 136-139.-Texella reyesi, new
Page 231 and 232: 1989 (W. Elliott, J. Reddell, and M
Page 233 and 234: Table 3.-Continued.# locality sex S
Page 235 and 236: mesoapical; patella, 2 mesal; tibia
Page 237 and 238: Figs. 162-165.-Texella gmbbsi, new
Page 239 and 240: Figs. 166-169.-Texella diplospina,
Page 241 and 242: Figs. 174-177.-Texella renkesae, ma
Page 243 and 244: Figs. 178-18\.-Teulla spinoperca, n
Page 245 and 246: Distribution.-Known only from Fayet
Page 247 and 248: Figs. 190-193.-Texellafendi, new sp
Page 249 and 250: CLASSIFICAnONTexellabifurcata group
Page 251 and 252: Chandler, D.S. 1992. The Pselaphida
Page 253: Key to Species1. Abdominal segments
Page 257 and 258: vertexal carinae, and the laterally
Page 259 and 260: is associated with rotten woods (Ch
Page 261 and 262: small ventral carina near base, pro
Page 263: Grigarick, A.A., and R.O. Schuster.
Page 266 and 267: Cicurifla (Cicurella) holsiflgeri G
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