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lateral carinae strong and extending to antennalbases, medial carina distinct and extending fromcervix over moderately swollen vertex to point anteriorto nude vertexal foveae, eyes with approximately45 facets; antennomere X large and globose,wider than XI, X with small ventral fovea near base.Pronotum sparsely punctate, median longitudinalsulcus extending to near apex; elytra with threebasal foveae and subhumeral fovea; sternite VIslightly flattened medially; second mesotarsomeresstraight.Females with coarse punctures only on antennaltubercles and posteriorly to near vertexal foveae,eyes with approximately 42 facets; lacking modificationsof head and antennae, sternite VI barelyrounded at middle.Male from Simons Squirm-Around Cave: antennae1.12, metafemora 0.79, metatibiae 0.76, metatarsi0.40.Specimens examined.-5: TEXAS: BurnetCounty: 1 male, Simons Squirm-Around Cave,XI-20-1990, J. Reddell, M. Reyes, underside ofrock loosely buried in soil (TMMC); 1 female,Road Side Sink No.1, XI-20-1990, J. Reddell,underside of rock loosely buried in soil (TMMC); 1male, Persimon Sink, 1-17-1991, J. Reddell, M.Reyes (DSC); 1 male, Simons Pretty Pit, 1-17-1991,J. Reddell (DSC); 1 female, Snake Pit Sink,11-8-1991, G. Veni (TMMC).Comments.-Batrisodes globosus is foundthroughout eastern North America, with isolatedrecords from Colorado, Washington, and Alberta. ItFigs. 8-17.-8-16, Balrisodes spp.: 8, B. reyesi, dorsal and right lateral view, aedeagus; 9, B. lexanus, head and anterior view,right antenna of male; 10, B. lexanus, lateral view, right antenna of female; II, B. lexanus, posterior view, right metatibia of male; 12,B. lexanus, dorsal view, aedeagus; 13, B. venyivi, head and anterior view, right antenna of male; 14, B. venyivi, dorsal and right lateralview, aedeagus; 15, B. globosus, head and anterior view, right antenna of male; 16, B. globosus, dorsal and right lateral view,aedeagus. Apical three segments of male antennae of Balrisodes spp. presented in ventral'view. 17, Texamaurops reddelli, head andanterior view, right antenna of female. Scale line equals 0.I.248
is associated with rotten woods (Chandler, 1987,and many collection records), and may be foundwith ants in the northern half of its range (Park,1960). Park (1960) noted one population that wasfound in a cave in Alabama.Male globosus may be separated from uncicomisby the more broadly rounded anterior margin of thevertex, and by the form of the apical segments of theantennae. Females lack coarse punctures between thevertexal fovea on the head, and also on the pronotum,while in uncicomis coarse punctures will befound in both areas.8. Texamaurops reddelli Barr and Steeves(Figs. 17-19)Texamaurops reddelli Barr and Steeves, 1963: 118.Type locality: Kretschmarr Cave, Travis Co.,Texas. Holotype female (FMNH).Description.-Length 2.72-3.08. Male with vertexsmoothly merging into elongate frons, areasparsely setate with prominent antennal tubercles,base of vertex with median and lateral carinae distinctbut not prominent, lateral carinae reachingapex of antennal tubercles, median extending topoint anterior to nude vertexal foveae, all antennomereselongate, X with large nude fovea on ventralsurface near base, antennomere swollen in basalhalf, eye area a rounded angulation with approximately6 granules that appear to be vestigial eyefacets.Pronotum with median longitudinal sulcus faint,extending to near pronotal apex; elytra with 2 basalfoveae and subhumeral fovea; metasternum atmiddle with group of long sparse setae; metatibiaelacking apical pencil of setae, sternite VI laterallywith rounded tumuli bearing long setae that aredirected laterally.Females with lateral margins of antennomere Xstraight, lacking ventral fovea, sternite VI broadlyrounded.Male from Stovepipe Cave: antennae 2.12,metafemora 1.56, metatibiae 1.72, metatarsi 0.80.Specimens examined.-6: TEXAS: TravisCounty: Holotype female, Kretschmarr Cave, 15 mi.NW Austin, III-2-1963, J. R. Reddell, D. McKenzie(FMNH); 1 female, same locality, 1968, R. W.Mitchell (FMNH); 1 female, Tooth Cave, V-31964, J. R. Reddell (TMMC); 1 female, samelocality, V-14-1966, J. R. Reddell (FMNH); 1female, Amber Cave, IV-8-1984, J. Reddell, M.Reyes (DSC); 1 male, Stovepipe Cave, X-25-1990,L. Sherrod, under rock lightly buried in silty clay insmal1 side room in total darkness (DSC).Comments.-Texamaurops reddelli has onlybeen found in four closely situated caves on theunderside of rocks lightly buried in silt in totaldarkness (James R. Reddel1, pers. comm.), and hasbeen placed on the US. List of Endangered Species(Chambers and Jahrsdoerfer, 1988). It has elongatelegs and antennae with reduced eyes, and is clearly atroglobite. The holotype and the single male aredifferent from all the other specimens in possessingonly two basal foveae on each elytron. The otherfour specimens, including one from the typelocality, have three equal foveae at the elytral base.AI1 other features appear to be similar and these fourspecimens are placed as reddelli.Barr and Steeves (1963) initially believed thatthis genus might be best placed in the Amauropsini,a European tribe whose members are all troglobites.Barr later (1974) figured the aedeagus of a purportedmale, which clearly indicated the placement of Texamauropsin the Batrisini since the morphology ofthe aedeagus in members of the two tribes is quitedifferent (Jeannel, 1948). The male specimen inquestion has proven to be a new species of Batrisodesconvergent with Texamaurops in appearance,but discovery of a true male of Texamauropsconfirms the placement in the Batrisini since theaedeagus is similar in form to those of members ofBatrisodes. The absence of the metatibial pencil ofsetae is shared by members of BatriasymmodesPark, a group with a number of troglobitic speciesin the eastern United States, and some members ofthis group also have a fovea on antennomere X.However, the aedeagal form of this genus is consistentlydifferent from members of Batriasymmodes,and based on the form of the aedeagus andantennal characters Texamaurops is probably bestconsidered a lineage derived from Batrisodes thathas lost the metatibial pencil of setae.Texamaurops reddelli is superficial1y similar totexanus by the greatly elongate antennae and legs, aswell as body size. However, reddelli possesses adistinct rounded angulate knob where the eyesshould be, and the metatibiae lack an apical pencilof setae.9. Cylindrarctus bicomis Chandler(Fig. 20)Cylindrarctus bicomis Chandler, 1988: 135. Typelocality: Texas (probably Bosque County). Holotypemale (USNM).249
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PREFACEThe present volume is the se
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TABLE OF CONTENTSHOLSINGER, JOHN R.
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the state of Coahuila in northern M
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Fig. 2.-Anesia welboumi, new specie
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\\. \ - -' ..........---~\ I// --..
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Mexiweckelia hardeni, new speciesFi
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2 sets of 1 or 2 setae each; dactyl
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AFig. 7.-Holsingerius smaragdinus,
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have as many setae on the inner pla
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Antenna 1 about 33 % length of body
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Of biogeographic interest for the h
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Bowman, T .E. 1992. Two subterranea
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A note by Scott Harden that accompa
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~~..~ ~~ I))\\ "-,'.=bFig. 3.-Speoc
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unarmed, except in pleopod 2, which
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Cokendolpher, LC., and l.R. Reddell
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lobes. The larger setae vary greatl
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the relationships of the order and
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zomids their absence is considered
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some species could be either split,
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have small pores over the surface o
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inhabits tropical deciduous forest
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huitvnolotitlensis from A. stygius
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8-10, figs. 5-7; Rowland, 1973c:136
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(in row) and one pair large posteri
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(0.34); tarsus 0.64 (0.58); total 4
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Abdomen: Tergite I with two pairs a
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setae near posterior margin. stemit
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setae, and ten ventral setae. Stemi
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1,980 m elev., 26 Dec. 1986 (T. Tre
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asitarsal-tarsal proportions: 15:4:
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and one pair setae at base of proce
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Cephalothorax: Propeltidium 1.66 mm
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Male adult unknown.Immature paratyp
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IV: trochanter 1.20 (1.10); femur 3
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Cokendolpher, 1.C. 1981. The order
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Gertsch, W.J. 1992. Distribution pa
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same families and genera, but also
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species from the United States and
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the posterior median pair short and
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great many species of North America
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Key to the Eyed Females1. Eight eye
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39111012Figs. 1-12.-Ventral and dor
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1419 23Figs. 13-24.-Ventral and dor
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27 293334Figs. 25-36.-Ventral and d
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Figs. 37-48.-Ventral and retrolater
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Cicurina blanco, new speciesFigs. 7
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Type-data.-Female holotype from ins
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Description.-Female holotype: Lengt
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lengths: first femur 2 rom, fourth
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Cicurina pablo, new speciesFigs. 10
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canal in nearly vertical posItion;
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Cicurina vespera, new speciesFigs.
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procurved canal across sac; connect
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Cicurina caverna, new speciesFigs.
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with sac of similar size set in obl
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February 1964 (J. Reddell, D. McKen
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County: Diamond Cave, 16 August 196
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Roth, V.D. 1992. A new and first tr
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LITERATURE CITEDBarr, T.C. 1963. Ec
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Muchmore, W.B. 1992. Cavernicolous
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Species of Aphrastochthonius are kn
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FAMILY NEOBISIIDAE CHAMBERLINGenus
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trochanter 2.5 (2.6), femur 5.55 (5
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ottom of entrance pit, Ogle Cave (O
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tactile seta on tibia and basitarsu
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Missouri may be conspecific (unpubl
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Trichobothriotaxy of chela generall
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Female (figures given first for all
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transverse furrows; eyespots not ev
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Reddell and W. Russell); I female f
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and Acuminochernes, along with the
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Chamberlin, J.C. 1946. The genera a
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(0.36); chela (without pedicel) 2.0
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although not recently studied is no
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and stylar outgrowths, present in s
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in Phalangodes (et al.) is thick an
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pairs) are found in six species: T.
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elated species IS probably best int
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legs than expected (2.6-3.2). This
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~ 3.3 are synapomorphic); all trogl
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TAXONOMYTEXELLA Goodnight and Goodn
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14. BK absent (Figs. 177, 180). SA
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Figs. 8-11.-Texella bijUrcata (Brig
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Description.-Total body length, 1.5
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male examined closely has fewer set
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Figs. 26-29.-Texella kokoweej, new
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apical region which loses the apica
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Figs. 38-41.-Texella shoshone, new
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Distribution.-Known only from the t
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Figs. 52-55.-Texella brevistyla, ne
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Texellajungi, new speciesFigs. 60-7
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Figs. 66-69.-Texellajungi, new spec
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than S; SA with laterobasal carina
Page 207 and 208: ~81//J/'/ ;'?/ ~~.........--~~I, II
Page 209 and 210: cylindrical, retina and cornea abse
Page 211 and 212: Figs. 93-96.-Texella cokendolpheri,
Page 213 and 214: SA with well developed prong and re
Page 215 and 216: Figs. 105-108.-Texelia mulaiki Good
Page 217 and 218: and McCarty Caves, which are known
Page 219 and 220: Color orange. Body of medium rugosi
Page 221 and 222: Notes.-The type locality was errone
Page 223 and 224: Figs. 128-131.-Taella reyesi, new s
Page 225 and 226: Figs. 136-139.-Texella reyesi, new
Page 231 and 232: 1989 (W. Elliott, J. Reddell, and M
Page 233 and 234: Table 3.-Continued.# locality sex S
Page 235 and 236: mesoapical; patella, 2 mesal; tibia
Page 237 and 238: Figs. 162-165.-Texella gmbbsi, new
Page 239 and 240: Figs. 166-169.-Texella diplospina,
Page 241 and 242: Figs. 174-177.-Texella renkesae, ma
Page 243 and 244: Figs. 178-18\.-Teulla spinoperca, n
Page 245 and 246: Distribution.-Known only from Fayet
Page 247 and 248: Figs. 190-193.-Texellafendi, new sp
Page 249 and 250: CLASSIFICAnONTexellabifurcata group
Page 251 and 252: Chandler, D.S. 1992. The Pselaphida
Page 253 and 254: Key to Species1. Abdominal segments
Page 255 and 256: stemite VI slightly impressed at ba
Page 257: vertexal carinae, and the laterally
Page 261 and 262: small ventral carina near base, pro
Page 263: Grigarick, A.A., and R.O. Schuster.
Page 266 and 267: Cicurifla (Cicurella) holsiflgeri G
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