12.07.2015 Views

searchable PDF - Association for Mexican Cave Studies

searchable PDF - Association for Mexican Cave Studies

searchable PDF - Association for Mexican Cave Studies

SHOW MORE
SHOW LESS
  • No tags were found...

You also want an ePaper? Increase the reach of your titles

YUMPU automatically turns print PDFs into web optimized ePapers that Google loves.

is here used in a more restricted sense to includeonly medium-sized structures (of about 20 /-Lm inbasal diameter) having rounded crowns. Included inthis definition are the paraocular anterior tubercles(AT), tubercles located on the posterior tergitemargins, ventral trochanteral tubercles, and bodytubercles scattered over dorsal and ventral surfaces.The smaller tuberculate structures, which constitutethe background texture of the phalangodid bodycuticle Cmicrotubercles' of Murphree, 1988), arereferred to as 'fine rugosity' (<strong>for</strong> microtubercles ofabout 5 /-Lm in diameter) or 'coarse rugosity' (<strong>for</strong>microtubercles of about 10 /-Lm in diameter). Forlarger structures, the terms 'eye mound' and'megaspine' (here used somewhat loosely to refer toboth the macroseta and the basal, cuticular process)are used to substitute 'eye tubercle' and 'palpalspine-bearing tubercle', respectively.Glans Nomenclature. The terminology used <strong>for</strong>the male genitalia is largely descriptive and novel.Most of the terms used previously <strong>for</strong> Calicina(Ubick and Briggs, 1989) are not repeated here inorder to avoid making premature assumptions abouthomology. Positional adjectives of glans structures,such as "the ventral surface of the stylus," refer to afully expanded glans where the stylus is apically directed.Tarsal Count. The tarsal count notation is abbreviated,as in our previous studies, with the numberof tarsomeres of legs I through IV listed seriallyand separated by a "-" (e.g., 3-5-5-5). However, inmany individuals of Texella the tarsal count is notbilaterally symmetrical and this variation is shownby a "I", with the value <strong>for</strong> the left leg given first.For example, "3-5-615-5" indicates asymmetry inleg III, the left leg having six tarsomeres and theright having five.Abbreviations:AS = apical spineAT = anterior tuberclesBF = basal fold of stylusBK = basal knob of glansBS = basal segment of glansDS = dorsal setae of VPPGO = genital operculumLIIISL = leg III scute lengthLS = lateral setae of VPPML = middle lobe of glansPOP = postopercular processPSL = parastylar lobe(s)PSL2 = secondary parastylar lobe(s)PTb = palpal tibiaS = stylusSA = stylar apophysisTC = tarsal countTrIV = trochanter of leg IVVC = ventral carina of stylusVP = ventral plateVPP = ventral plate prongVS = ventral setae of VPPMORPHOLOGYOf the 29 characters used in this analysis the vastmajority (20) are from the male genitalia. Themorphologically simpler female genitalia werecomparably poorer in apparent characters (2). Inaddition, secondary sexual characters (3) and somesomatic characters (4) were found useful.Male genitalia.-(Figs. 6-7). The penis ofTexella consists of an inflatable basal sac, a sclerotizedtruncus, and a distal glans. The ventroapicalextension of the truncus, the ventral plate (VP), isdeeply bifurcate. The resulting prongs (VPP) arelateral in position and widely separated, permittingthe ventral protrusion of the glans. The prongs bearsetae arranged on the dorsal, lateral, and ventralsurfaces (OS, LS, and VS, respectively). Also onthe lateral surface, and somewhat distad of thelateral setae, is a stout cuticular outgrowth, theapical spine (AS).The glans of Texella is robust and appearssigmoid in lateral view. The basal segment (BS) islarge, often approaching the apex of the VPP, and,in roughly half of the species, bears on itsventroapical margin a pointed extension, the basalknob (BK). The middle segment is split apically to<strong>for</strong>m two lobes, the parastylar lobes (PSL), and, insome species, an additional pair of lobes, thesecondary parastylar lobes (PSL2). The middle lobe(ML) is a thin flap which separates the middlesegment of the glans from the apical segment, thestylus (S). The stylus is either simple or has a basalenlargement (the basal fold, BF), a lateral process(the stylar apophysis, SA), andlor a membranousventral carina (VC).Of all the Nearctic phalangodid genera, the penisin Banksula (Fig. 5) is most similar to that of TexellaoOne obvious difference is the placement of theVPP, which are relatively contiguous and positionedon the ventral margin of the truncus. Also, theglans is considerably simpler than in Texella: theglans has only one axis of rotation, the basalsegment (BS) is reduced, and the apical elementslack accessory structures (except <strong>for</strong> a pair of PSL158

Hooray! Your file is uploaded and ready to be published.

Saved successfully!

Ooh no, something went wrong!