is here used in a more restricted sense to includeonly medium-sized structures (of about 20 /-Lm inbasal diameter) having rounded crowns. Included inthis definition are the paraocular anterior tubercles(AT), tubercles located on the posterior tergitemargins, ventral trochanteral tubercles, and bodytubercles scattered over dorsal and ventral surfaces.The smaller tuberculate structures, which constitutethe background texture of the phalangodid bodycuticle Cmicrotubercles' of Murphree, 1988), arereferred to as 'fine rugosity' (<strong>for</strong> microtubercles ofabout 5 /-Lm in diameter) or 'coarse rugosity' (<strong>for</strong>microtubercles of about 10 /-Lm in diameter). Forlarger structures, the terms 'eye mound' and'megaspine' (here used somewhat loosely to refer toboth the macroseta and the basal, cuticular process)are used to substitute 'eye tubercle' and 'palpalspine-bearing tubercle', respectively.Glans Nomenclature. The terminology used <strong>for</strong>the male genitalia is largely descriptive and novel.Most of the terms used previously <strong>for</strong> Calicina(Ubick and Briggs, 1989) are not repeated here inorder to avoid making premature assumptions abouthomology. Positional adjectives of glans structures,such as "the ventral surface of the stylus," refer to afully expanded glans where the stylus is apically directed.Tarsal Count. The tarsal count notation is abbreviated,as in our previous studies, with the numberof tarsomeres of legs I through IV listed seriallyand separated by a "-" (e.g., 3-5-5-5). However, inmany individuals of Texella the tarsal count is notbilaterally symmetrical and this variation is shownby a "I", with the value <strong>for</strong> the left leg given first.For example, "3-5-615-5" indicates asymmetry inleg III, the left leg having six tarsomeres and theright having five.Abbreviations:AS = apical spineAT = anterior tuberclesBF = basal fold of stylusBK = basal knob of glansBS = basal segment of glansDS = dorsal setae of VPPGO = genital operculumLIIISL = leg III scute lengthLS = lateral setae of VPPML = middle lobe of glansPOP = postopercular processPSL = parastylar lobe(s)PSL2 = secondary parastylar lobe(s)PTb = palpal tibiaS = stylusSA = stylar apophysisTC = tarsal countTrIV = trochanter of leg IVVC = ventral carina of stylusVP = ventral plateVPP = ventral plate prongVS = ventral setae of VPPMORPHOLOGYOf the 29 characters used in this analysis the vastmajority (20) are from the male genitalia. Themorphologically simpler female genitalia werecomparably poorer in apparent characters (2). Inaddition, secondary sexual characters (3) and somesomatic characters (4) were found useful.Male genitalia.-(Figs. 6-7). The penis ofTexella consists of an inflatable basal sac, a sclerotizedtruncus, and a distal glans. The ventroapicalextension of the truncus, the ventral plate (VP), isdeeply bifurcate. The resulting prongs (VPP) arelateral in position and widely separated, permittingthe ventral protrusion of the glans. The prongs bearsetae arranged on the dorsal, lateral, and ventralsurfaces (OS, LS, and VS, respectively). Also onthe lateral surface, and somewhat distad of thelateral setae, is a stout cuticular outgrowth, theapical spine (AS).The glans of Texella is robust and appearssigmoid in lateral view. The basal segment (BS) islarge, often approaching the apex of the VPP, and,in roughly half of the species, bears on itsventroapical margin a pointed extension, the basalknob (BK). The middle segment is split apically to<strong>for</strong>m two lobes, the parastylar lobes (PSL), and, insome species, an additional pair of lobes, thesecondary parastylar lobes (PSL2). The middle lobe(ML) is a thin flap which separates the middlesegment of the glans from the apical segment, thestylus (S). The stylus is either simple or has a basalenlargement (the basal fold, BF), a lateral process(the stylar apophysis, SA), andlor a membranousventral carina (VC).Of all the Nearctic phalangodid genera, the penisin Banksula (Fig. 5) is most similar to that of TexellaoOne obvious difference is the placement of theVPP, which are relatively contiguous and positionedon the ventral margin of the truncus. Also, theglans is considerably simpler than in Texella: theglans has only one axis of rotation, the basalsegment (BS) is reduced, and the apical elementslack accessory structures (except <strong>for</strong> a pair of PSL158
and stylar outgrowths, present in some species,which mayor may not be homologs to the SA).Function: The glans of Texella is of the foldedtype. In its unexpanded <strong>for</strong>m, the stylus and the(deflated) middle segment are pressed against the(deflated) basal segment. During expansion, assimulated by treatment in KOH, the membranousportions of the glans inflate and the distal elementsrotate along a dorsoapical arc. There are two axesof rotation: the primary, at the junction of the basaland middle segments, and the secondary at the baseof the stylus. With complete expansion the styluspoints apically (roughly 180 0 from the unfoldedcondition) and the entire glans protrudes ventrallybeyond the VPP. At this stage, the stylus is typicallyseparated from the parastylar lobes, which abductand (in some species) inflate and contort topermit passage of the stylus.It is worth noting that, in virtually all instancesof unexpanded and partially expanded glandes observed,the parastylar lobes are in intimate contactwith the stylus. It appears that the PSL lock the S inposition. Three types of these apparent lock mechanismswere found: 1) PSL-S. In this, the mostcommon type of lock, the PSL adduct, pinching theS (Figs. 68, 87, 96). 2) PSL-SA. In this type,found only in species of the spinoperca infragroup,the PSL grip the attenuated prongs of the SA (Fig.180). 3) SA-PSL. In this type, found only in thereddelli infragroup, the clip-like SA grip the basalportions of the PSL (Figs. 120, 145).Female genitalia.-The ovipositor in Texella, asin other Nearctic Phalangodidae, is structurallyrelatively simple (Fig. 18). The cuticle is conspicuouslywrinkled (with longitudinal folds on the ventraland dorsal surfaces and transverse folds alongthe lateral surfaces) and sculptured (with minutetubercles and spines, often arranged in linear serrations).The apical surface bears a pair of cuticularprojections (= apical teeth) and 7 (rarely 6) pairs ofsetae, arranged as in Fig. 20.The ovipositor of BanksuLa resembles that ofTexella but lacks apical teeth (at least in the threespecies closely examined: B. grahami Briggs,meLones Briggs, and rudoLphi Briggs and Ubick).One noteworthy observation is the presence of ahyperexpanded ovipositor (Figs. 152-155) whichdiffers from the typical <strong>for</strong>m in having an additionalapical segment terminating in four lobes. Thishyperexpanded state, most likely being the conditionassumed during egg laying, was also observed by usin a few specimens of Bishopella sp. and"SitaLcina" cockerelli Goodnight and Goodnight butwas not recorded by Martens, Hoheisel, and Gatze(1981) in their study of opilionid ovipositor morphology.However, thanks to the detailed drawingsVSVPP,..ASDSVS, /LS , ,LS, , ,,, .5 6 7Figs. 5-7.-Phalangodid penes, dorsolateral views: 5, Bank.mla galilei Briggs; 6, Texella bifurcara (Briggs); 7. Texella reyesi,new species. Styli shaded. Abbreviations: AS = apical spine, BF = basal fold of stylus, BK = basal knob of glans, BS = basalsegment of glans, DS = dorsal setae, LS = lateral setae, ML = middle lobe of glans, PSL = parastylar lobes, PSL2 = secondaryparastylar lobes, SA = stylar apophysis, VC = ventral carina of stylus, VPP = ventral plate prong, VS = ventral setae.159
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PREFACEThe present volume is the se
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TABLE OF CONTENTSHOLSINGER, JOHN R.
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the state of Coahuila in northern M
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Fig. 2.-Anesia welboumi, new specie
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\\. \ - -' ..........---~\ I// --..
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Mexiweckelia hardeni, new speciesFi
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2 sets of 1 or 2 setae each; dactyl
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AFig. 7.-Holsingerius smaragdinus,
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have as many setae on the inner pla
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Antenna 1 about 33 % length of body
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Of biogeographic interest for the h
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Bowman, T .E. 1992. Two subterranea
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A note by Scott Harden that accompa
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~~..~ ~~ I))\\ "-,'.=bFig. 3.-Speoc
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unarmed, except in pleopod 2, which
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Cokendolpher, LC., and l.R. Reddell
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lobes. The larger setae vary greatl
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the relationships of the order and
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zomids their absence is considered
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some species could be either split,
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have small pores over the surface o
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inhabits tropical deciduous forest
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huitvnolotitlensis from A. stygius
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8-10, figs. 5-7; Rowland, 1973c:136
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(in row) and one pair large posteri
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(0.34); tarsus 0.64 (0.58); total 4
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Abdomen: Tergite I with two pairs a
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setae near posterior margin. stemit
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setae, and ten ventral setae. Stemi
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1,980 m elev., 26 Dec. 1986 (T. Tre
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asitarsal-tarsal proportions: 15:4:
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and one pair setae at base of proce
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Cephalothorax: Propeltidium 1.66 mm
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Male adult unknown.Immature paratyp
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IV: trochanter 1.20 (1.10); femur 3
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Cokendolpher, 1.C. 1981. The order
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Gertsch, W.J. 1992. Distribution pa
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same families and genera, but also
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species from the United States and
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the posterior median pair short and
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great many species of North America
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Key to the Eyed Females1. Eight eye
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39111012Figs. 1-12.-Ventral and dor
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1419 23Figs. 13-24.-Ventral and dor
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27 293334Figs. 25-36.-Ventral and d
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Figs. 37-48.-Ventral and retrolater
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Cicurina blanco, new speciesFigs. 7
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Type-data.-Female holotype from ins
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Description.-Female holotype: Lengt
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Description.-Female holotype: Lengt
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lengths: first femur 2 rom, fourth
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Cicurina pablo, new speciesFigs. 10
- Page 117 and 118: Description.-Female holotype: Lengt
- Page 119 and 120: canal in nearly vertical posItion;
- Page 121 and 122: Cicurina vespera, new speciesFigs.
- Page 123 and 124: procurved canal across sac; connect
- Page 125 and 126: Cicurina caverna, new speciesFigs.
- Page 127 and 128: with sac of similar size set in obl
- Page 129 and 130: February 1964 (J. Reddell, D. McKen
- Page 131 and 132: County: Diamond Cave, 16 August 196
- Page 133 and 134: Roth, V.D. 1992. A new and first tr
- Page 135 and 136: LITERATURE CITEDBarr, T.C. 1963. Ec
- Page 137 and 138: Muchmore, W.B. 1992. Cavernicolous
- Page 139 and 140: Species of Aphrastochthonius are kn
- Page 141 and 142: FAMILY NEOBISIIDAE CHAMBERLINGenus
- Page 143 and 144: trochanter 2.5 (2.6), femur 5.55 (5
- Page 145 and 146: ottom of entrance pit, Ogle Cave (O
- Page 147 and 148: tactile seta on tibia and basitarsu
- Page 149 and 150: Missouri may be conspecific (unpubl
- Page 151 and 152: Trichobothriotaxy of chela generall
- Page 153 and 154: Female (figures given first for all
- Page 155 and 156: transverse furrows; eyespots not ev
- Page 157 and 158: Reddell and W. Russell); I female f
- Page 159 and 160: and Acuminochernes, along with the
- Page 161 and 162: Chamberlin, J.C. 1946. The genera a
- Page 163: (0.36); chela (without pedicel) 2.0
- Page 167: although not recently studied is no
- Page 171 and 172: in Phalangodes (et al.) is thick an
- Page 173 and 174: pairs) are found in six species: T.
- Page 175 and 176: elated species IS probably best int
- Page 177 and 178: legs than expected (2.6-3.2). This
- Page 179 and 180: ~ 3.3 are synapomorphic); all trogl
- Page 181 and 182: TAXONOMYTEXELLA Goodnight and Goodn
- Page 183 and 184: 14. BK absent (Figs. 177, 180). SA
- Page 185 and 186: Figs. 8-11.-Texella bijUrcata (Brig
- Page 187 and 188: Description.-Total body length, 1.5
- Page 189 and 190: male examined closely has fewer set
- Page 191 and 192: Figs. 26-29.-Texella kokoweej, new
- Page 193 and 194: apical region which loses the apica
- Page 195 and 196: Figs. 38-41.-Texella shoshone, new
- Page 197 and 198: Distribution.-Known only from the t
- Page 199 and 200: Figs. 52-55.-Texella brevistyla, ne
- Page 201 and 202: Texellajungi, new speciesFigs. 60-7
- Page 203 and 204: Figs. 66-69.-Texellajungi, new spec
- Page 205 and 206: than S; SA with laterobasal carina
- Page 207 and 208: ~81//J/'/ ;'?/ ~~.........--~~I, II
- Page 209 and 210: cylindrical, retina and cornea abse
- Page 211 and 212: Figs. 93-96.-Texella cokendolpheri,
- Page 213 and 214: SA with well developed prong and re
- Page 215 and 216: Figs. 105-108.-Texelia mulaiki Good
- Page 217 and 218: and McCarty Caves, which are known
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Color orange. Body of medium rugosi
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Notes.-The type locality was errone
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Figs. 128-131.-Taella reyesi, new s
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Figs. 136-139.-Texella reyesi, new
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Figs. 144-147.-Texella reyesi, new
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Figs. 152-155.-Texella reyesi, new
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1989 (W. Elliott, J. Reddell, and M
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Table 3.-Continued.# locality sex S
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mesoapical; patella, 2 mesal; tibia
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Figs. 162-165.-Texella gmbbsi, new
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Figs. 166-169.-Texella diplospina,
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Figs. 174-177.-Texella renkesae, ma
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Figs. 178-18\.-Teulla spinoperca, n
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Distribution.-Known only from Fayet
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Figs. 190-193.-Texellafendi, new sp
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CLASSIFICAnONTexellabifurcata group
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Chandler, D.S. 1992. The Pselaphida
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Key to Species1. Abdominal segments
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stemite VI slightly impressed at ba
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vertexal carinae, and the laterally
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is associated with rotten woods (Ch
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small ventral carina near base, pro
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Grigarick, A.A., and R.O. Schuster.
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Cicurifla (Cicurella) holsiflgeri G