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searchable PDF - Association for Mexican Cave Studies

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and stylar outgrowths, present in some species,which mayor may not be homologs to the SA).Function: The glans of Texella is of the foldedtype. In its unexpanded <strong>for</strong>m, the stylus and the(deflated) middle segment are pressed against the(deflated) basal segment. During expansion, assimulated by treatment in KOH, the membranousportions of the glans inflate and the distal elementsrotate along a dorsoapical arc. There are two axesof rotation: the primary, at the junction of the basaland middle segments, and the secondary at the baseof the stylus. With complete expansion the styluspoints apically (roughly 180 0 from the unfoldedcondition) and the entire glans protrudes ventrallybeyond the VPP. At this stage, the stylus is typicallyseparated from the parastylar lobes, which abductand (in some species) inflate and contort topermit passage of the stylus.It is worth noting that, in virtually all instancesof unexpanded and partially expanded glandes observed,the parastylar lobes are in intimate contactwith the stylus. It appears that the PSL lock the S inposition. Three types of these apparent lock mechanismswere found: 1) PSL-S. In this, the mostcommon type of lock, the PSL adduct, pinching theS (Figs. 68, 87, 96). 2) PSL-SA. In this type,found only in species of the spinoperca infragroup,the PSL grip the attenuated prongs of the SA (Fig.180). 3) SA-PSL. In this type, found only in thereddelli infragroup, the clip-like SA grip the basalportions of the PSL (Figs. 120, 145).Female genitalia.-The ovipositor in Texella, asin other Nearctic Phalangodidae, is structurallyrelatively simple (Fig. 18). The cuticle is conspicuouslywrinkled (with longitudinal folds on the ventraland dorsal surfaces and transverse folds alongthe lateral surfaces) and sculptured (with minutetubercles and spines, often arranged in linear serrations).The apical surface bears a pair of cuticularprojections (= apical teeth) and 7 (rarely 6) pairs ofsetae, arranged as in Fig. 20.The ovipositor of BanksuLa resembles that ofTexella but lacks apical teeth (at least in the threespecies closely examined: B. grahami Briggs,meLones Briggs, and rudoLphi Briggs and Ubick).One noteworthy observation is the presence of ahyperexpanded ovipositor (Figs. 152-155) whichdiffers from the typical <strong>for</strong>m in having an additionalapical segment terminating in four lobes. Thishyperexpanded state, most likely being the conditionassumed during egg laying, was also observed by usin a few specimens of Bishopella sp. and"SitaLcina" cockerelli Goodnight and Goodnight butwas not recorded by Martens, Hoheisel, and Gatze(1981) in their study of opilionid ovipositor morphology.However, thanks to the detailed drawingsVSVPP,..ASDSVS, /LS , ,LS, , ,,, .5 6 7Figs. 5-7.-Phalangodid penes, dorsolateral views: 5, Bank.mla galilei Briggs; 6, Texella bifurcara (Briggs); 7. Texella reyesi,new species. Styli shaded. Abbreviations: AS = apical spine, BF = basal fold of stylus, BK = basal knob of glans, BS = basalsegment of glans, DS = dorsal setae, LS = lateral setae, ML = middle lobe of glans, PSL = parastylar lobes, PSL2 = secondaryparastylar lobes, SA = stylar apophysis, VC = ventral carina of stylus, VPP = ventral plate prong, VS = ventral setae.159

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