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searchable PDF - Association for Mexican Cave Studies

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Distribution.-Known only from the type locality(Map 1).The mulaiki GroupDiagnosis.-The following combination of malecharacters distinguishes the members of this group:POP present (secondarily lost in T. mulaiki,hardeni, bilobata, !eflai, and homi), glans peniswith BK (lost in T. brevistyla and the spiflopercainfragroup), and stylus with BF (small in T.brevistyla) (Figs. 7, 132, 133). No characters havebeen found which distinguish females.Range.-Known from southern New Mexico tocentral Texas (Maps 2-4).The brevistyla SubgroupDiagnosis.-The combination of S shorter thanPSL, and the absence of a SA (Figs. 52-55, 66-69,74-76) distinguishes males of these species from allother Texella.Range.-Known from western central Texas(Maps 3,4).Texella brevistyla, new speciesFigs. 48-59, 76.Diagnosis.-Males of this species are distinguishedfrom other Texella by the following combinationof characters: VPP apically convergent; PSLlong, attenuated, spiraled; S short, tube-like; BFpresent; SA absent; BK absent (Figs. 52-55, 76).Type.-Male holotype from Crom <strong>Cave</strong>, UvaldeCounty, Texas (Summer 1983; R. Waters), depositedin CAS.Etymology.- The speci fie name refers to theshort stylus characteristic of this species.Description.-Total body length, 1.56-2.18.Scute length, 1.18-1.44. Leg II length, 3.54-4.03.Leg II/Scute length, 2.58-3.17. (N = 11).Color orange to brownish orange. Body ofmedium rugosity; tubercles on eye mound, pars thoracica,tergite margins, and coxae. Carapace with 8pairs of AT arranged in two rows. Eye moundbroadly conical, eyes well developed, cervicalgroove prominent. Palpal megaspines: trochanter,1 ventral; femur, 1 mesoapical; patella, 2 mesal;tibia, 3 mesal. Tarsal count: 3-5-4-5.Male (holotype): Total body length, 1.85. Scutelength, 1.38; width, 1.44. Eye mound length, 0.41;width, 0.41. Leg II length, 3.56. TrIV spur robust,apically bent; length, 0.51. POP length, 0.26.Penis (Figs. 52-55, 76): VPP apically converging;with 2 dorsal, 10 lateral, and 4 ventral setae;AS short, curved. Glans: BK absent; ML present,of moderate sire; PSL elongate, tapering apically,spiraled. S short, tubular; BF present, reduced; SAabsent.Female (paratopotype): Total body length, 1.64.Scute length, 1.28; width, 1.33. Eye mound length,0.32; width, 0.31. Leg II length, 3.54. TrIV withtwo ventral tubercles.Ovipositor (Figs. 56, 58, 59): cuticle intricatelyfolded; dorsal surface sparsely set with microserrations,ventral smooth; 1 pair of small apical teethpresent; setal pattern: 1 pair dorsal, 4 pairs lateral, 1pair ventral.Variation.-The cervical groove is more prominentin specimens from Crom <strong>Cave</strong>. One male(SEM) has a short TrIV spur (L = 0.35) and astrongly reduced POP (L = 0.05) (Figs. 50, 51); inthe remaining males (6) these structures are uni<strong>for</strong>mlylarge (about 0.50 and 0.30 respectively).The aberrant male is distinctly smaller than theothers (scute length = 1.18 compared to 1.31-1.44<strong>for</strong> the others). One male from BFS <strong>Cave</strong> (SEM)lacks an apical spine (Figs. 52-55) which, like thetip of one prong, may be broken. The ovipositor ofthe female from BFS <strong>Cave</strong> (SEM) lacks a pair ofapical teeth (Figs. 56, 58) which are present,although small, in all three females from Crom<strong>Cave</strong>.Note.-The PSL are spiral-shaped and enclosethe S in the unexpanded and partially expandedglans. With more complete expansion the PSL separateand become somewhat unspiraled (as in Fig.53).Natural History.-One female (Crom <strong>Cave</strong>) haswhat appears to be two large eggs (one visiblethrough the somewhat transparent sternites and thesecond revealed through dissection). The presumedeggs are quite large (0.36 x 0.46), much larger thanthe genital opening (0.18 x 0.23), and occupy muchof the abdominal cavity. We have seen specimens ofanother laniatorid harvestman, Hoplobunus species(Gonyleptoidea), from a nearby cave, Frio Queen<strong>Cave</strong>.Other Material Examined (Paratypes).- UNIT­ED STATES: Texas: Uvalde Co.: Crom <strong>Cave</strong>,Summer 1983 (R. Waters, TMM, CDU), 3 females;BFS <strong>Cave</strong>, 8 June 1989 (A. Grubbs, "A.c.", and R. Waters, TMM, CAS, CDU), 6 males, 1female.Distribution.-Known only from Crom and BFScaves, Uvalde County, Texas (Maps 2-4).187

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