and in several leaf litters, and has been previouslyrecorded from a cave (Park, 1960: 1(0). A specimenof Cylindraretus bicomis Chandler was found in acave, but is presumed to be associated with leaf litters,based on the common association of the otherspecies in the genus. Trimioarcus musamator n. sp.has reduced eyes, but is most likely an inhabitant ofrodent nests or deep leaf litter since no other speciesin its tribe, the Euplectini, is known to be cavernicolous.In summary, five species are troglobites,one a myrmecophile, one a troglophile, and theother five species are probably only found in cavesnear their entrances or in litter piles washed in orbrought in by rodents.All measurements are in millimeters. The collectionswhere specimens are deposited are indicated bythe following codons: ANSP, Academy of NaturalSciences, Philadelphia; DSC, collection of author;FMNH, Field Museum of Natural History, Chicago;JAW, John A. Wagner collection, Evanston,Illinois; TMMC, Texas Memorial Museum Collection,Austin; USNM, National Museum of NaturalHistory, Washington, D.C. The types of all previouslydescribed species were examined, and theholotypes of species described here are to be placedin the Field Museum of Natural History, Chicago.Illustrations and descriptions were based on wholeand disarticulated specimens in temporary mountson slides, and checked against whole specimensmounted on points.SYSTEMATICS1. Batrisodes (Babnormodes) uncicomis (Casey)(Figs. 1-2)Batrisus uncicomis Casey, 1897:576. Type locality:New York City, New York. Male holotype(USNM).Batrisodes schneiderensis Park, 1960:75. Typelocality: Schneider <strong>Cave</strong>, Texas. Holotype female(FMNH). NEW SYNONYMY.Description.-Length 2.32-2.64. Males withhead transversely excavate anterior to antennalbases, frons lacking projections, largely covereddorsally by rounded angulate projection of vertex,dense setae projecting ventrally from anteriormargin of vertex between antennal bases, vertexcoarsely punctate especially on anterior half, lateralcarinae strong basally but fading be<strong>for</strong>e reachingantennal bases, median carina distinct but not strongand extending anteriorly to line between nudevertexal foveae, eyes with about 32 facets; firstantennomere (scape) not enlarged but conspicuouslypunctate on antero-ventral margin, antennomere IXprotruding laterally as a lamina, IX less than twiceas wide as long, X with large fovea occupying overhalf of ventral surface, XI with large ventraltubercle at base.Pronotum with large scattered punctures particularlyanteriorly, median longitudinal sulcus extendingto near apex; elytra with three basal foveae andsubhumeral fovea; sternite VI vaguely impressedlongitudinally; protibiae flattened near apex anddensely setate on medial margin, mesotibiaeexcavate on mesal margin in apical third, thick setaealong posterior edge of excavation, secondmesotarsomere sinuate ventrally near base.Females with coarse punctures on head and pronotum;eyes with about 17 facets, lacking modificationsof head, antennae, and legs; sternite VIrounded.Male from Schneider Ranch <strong>Cave</strong>: antennae 1.10,metafemora 0.84, metatibiae 0.80, metatarsi 0.37.Specimens examined.-43: TEXAS: BurnetCounty: Resurrection Well, VII- 2-1989, M. Grimm(1). Hays County: Anaqua <strong>Cave</strong>, VI-2I-I985, A.Grubbs (2); Ezell's <strong>Cave</strong>, 1-16-1979, J. C. Davis,cave area (2); Fern <strong>Cave</strong>, V-26-I989, A. Grubbs, J.Reddell, M. Reyes (5); Ladder <strong>Cave</strong>, IX-2-I989, D.Ubick, S. Fend, S. Renkes (1); McGlothlin Sink,V-26-I989, A. Grubbs, J. Reddell, M. Reyes (1).Carnal County: Klar's <strong>Cave</strong>, III-I2-I988, S. Spence,G. Veni (1). Kendall County: The Crack,V-28-I990, D. Pate (1); Schwarz <strong>Cave</strong>,IX-12-1987, J. Ivy, G. Veni (I); Schneider <strong>Cave</strong>,VIII-30-1959, T. C. Barr, Jr., H. M. Koepke (typeof schneiderensis); same locality, IX-8-1963, J.Reddell, D. McKenzie (4). Llano County:Enchanted Rock <strong>Cave</strong>, IV-I4-1985, J. Reddell, M.Reyes (1); Freshman Mountain <strong>Cave</strong>, IV-9-1989,W. Elliott, J. Reddell, M. Reyes (2). TravisCounty: Bandit <strong>Cave</strong>, V-I7-1965, T. C. Barr (I);<strong>Cave</strong> X, III-30-1974, W. Elliott, W. Russell, S. &R. Fieseler, C. Rogers (1); Ireland's <strong>Cave</strong>, 1-231989, J. Reddell, M. Reyes, E. Grimm, M. Grimm(10); laCrosse <strong>Cave</strong>, V-8-1990, J. Reddell, M.Reyes (2); Pickle Pit, V-2I-1990, J. Reddell, M.Reyes, L. Sherrod (2). Williamson County: BeckRanch <strong>Cave</strong> (=Beck's <strong>Cave</strong>), III-24-1989, J. Reddell,M. Reyes (1); same locality, V-16-I965, T. C.Barr (3). Specimens in collections of: DSC, FMNH,JAW, and TMMC.Comments.-Batrisodes uncicornis is a widespreadspecies whose range extends from Massachusettsand Florida to Texas. Habitat in<strong>for</strong>mation hasbeen associated with only a few specimens, and242
Key to Species1. Abdominal segments II-IV with acute lateral margins 2Abdominal segments II-IV lacking lateral margins, abdomenround in cross section (Fig. 1) (Batrisinae, Batrisini) 52.(1) Last segment of maxillary palpi laterally angulate, previous twosegments with spine on outer face (Fig. 21) (Pselaphinae,Tmesiphorini10. Tmesiphorus costalisApical segment of maxillary palpi elongate, lacking lateralangulation or spines on previous segments (Figs. 20, 22) 33.(2) Last segment of maxillary palpi elongate, penultimate segmentangulate on mesal margin (Fig. 20) (Goniacerinae, Tychini9. Cylindrarctus bicornisLast segment of maxillary palpi enlarged, penultimate segmentmuch smaller (Figs. 22, 24) 44.(3) Antennal club of last three segments, penultimate antennomerelonger than wide, last segment of maxillary palpus withlongitudinal groove on inner margin (Fig. 22) (Pselaphinae, Tyrini) 11. Hamotus electraeAntennal club of last two segments, penultimate antennomeremuch wider than long, last segment of maxillary pal pus lackingany groove (Fig. 24) (Euplectinae, Euplectini)12. Trimioarcus musamator, n. sp.5.(1) Apex of metatibiae lacking elongate pencil of appressed setae(Fig. 18); all antennomeres more than twice as long as wide(Fig. 17)8. Texamaurops reddelliApex of metatibiae with elongate pencil of appressed setae (Fig. 11);with antennomere VIII distinctly less than twice as long aswide except texanus with VIII twice as long as wide (Figs. 3, 9)(Batrisodes) 66.(5) Eyes distinct, with 10 to 50 facets grouped together. 7Eyes apparently absent, possibly up to 10 disassociated palegranules in area where eyes should be 97.(6) Vertex of head and anterior portion of pronotum coarselypunctate; males with anterior margin of vertex angulate (Fig. 1);females with vertex coarsely punctate1. Batrisodes uncicornisBasal half of vertex and anterior portion of pronotum not coarselypunctate; males with anterior margin of vertex truncate tobroadly rounded (Figs. 3, 15); females with vertex at mostcoarsely punctate on antennal tubercles, area between vertexalfoveae smooth or sparsely granulate 88.(7) Males with anterior margin of vertex broadly rounded,penultimate antennomere with small fovea at base (Fig. 15);females with large eyes of more than 40 facetsMales with anterior margin of vertex broadly truncate,penultimate antennomere lacking basal fovea (Fig. 3); femaleswith small eyes of approximately 15 facets7. Batrisodes globosus2. Batrisodes clypeonotus243
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PREFACEThe present volume is the se
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TABLE OF CONTENTSHOLSINGER, JOHN R.
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the state of Coahuila in northern M
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Fig. 2.-Anesia welboumi, new specie
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\\. \ - -' ..........---~\ I// --..
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Mexiweckelia hardeni, new speciesFi
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2 sets of 1 or 2 setae each; dactyl
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AFig. 7.-Holsingerius smaragdinus,
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have as many setae on the inner pla
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Antenna 1 about 33 % length of body
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Of biogeographic interest for the h
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Bowman, T .E. 1992. Two subterranea
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A note by Scott Harden that accompa
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~~..~ ~~ I))\\ "-,'.=bFig. 3.-Speoc
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unarmed, except in pleopod 2, which
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Cokendolpher, LC., and l.R. Reddell
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lobes. The larger setae vary greatl
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the relationships of the order and
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zomids their absence is considered
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some species could be either split,
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have small pores over the surface o
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inhabits tropical deciduous forest
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huitvnolotitlensis from A. stygius
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8-10, figs. 5-7; Rowland, 1973c:136
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(in row) and one pair large posteri
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(0.34); tarsus 0.64 (0.58); total 4
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Abdomen: Tergite I with two pairs a
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setae near posterior margin. stemit
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setae, and ten ventral setae. Stemi
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1,980 m elev., 26 Dec. 1986 (T. Tre
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asitarsal-tarsal proportions: 15:4:
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and one pair setae at base of proce
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Cephalothorax: Propeltidium 1.66 mm
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Male adult unknown.Immature paratyp
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IV: trochanter 1.20 (1.10); femur 3
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Cokendolpher, 1.C. 1981. The order
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Gertsch, W.J. 1992. Distribution pa
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same families and genera, but also
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species from the United States and
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the posterior median pair short and
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great many species of North America
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Key to the Eyed Females1. Eight eye
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39111012Figs. 1-12.-Ventral and dor
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1419 23Figs. 13-24.-Ventral and dor
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27 293334Figs. 25-36.-Ventral and d
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Figs. 37-48.-Ventral and retrolater
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Cicurina blanco, new speciesFigs. 7
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Type-data.-Female holotype from ins
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Description.-Female holotype: Lengt
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Description.-Female holotype: Lengt
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lengths: first femur 2 rom, fourth
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Cicurina pablo, new speciesFigs. 10
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Description.-Female holotype: Lengt
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canal in nearly vertical posItion;
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Cicurina vespera, new speciesFigs.
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procurved canal across sac; connect
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Cicurina caverna, new speciesFigs.
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with sac of similar size set in obl
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February 1964 (J. Reddell, D. McKen
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County: Diamond Cave, 16 August 196
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Roth, V.D. 1992. A new and first tr
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LITERATURE CITEDBarr, T.C. 1963. Ec
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Muchmore, W.B. 1992. Cavernicolous
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Species of Aphrastochthonius are kn
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FAMILY NEOBISIIDAE CHAMBERLINGenus
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trochanter 2.5 (2.6), femur 5.55 (5
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ottom of entrance pit, Ogle Cave (O
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tactile seta on tibia and basitarsu
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Missouri may be conspecific (unpubl
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Trichobothriotaxy of chela generall
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Female (figures given first for all
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transverse furrows; eyespots not ev
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Reddell and W. Russell); I female f
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and Acuminochernes, along with the
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Chamberlin, J.C. 1946. The genera a
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(0.36); chela (without pedicel) 2.0
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although not recently studied is no
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and stylar outgrowths, present in s
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in Phalangodes (et al.) is thick an
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pairs) are found in six species: T.
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elated species IS probably best int
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legs than expected (2.6-3.2). This
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~ 3.3 are synapomorphic); all trogl
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TAXONOMYTEXELLA Goodnight and Goodn
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14. BK absent (Figs. 177, 180). SA
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Figs. 8-11.-Texella bijUrcata (Brig
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Description.-Total body length, 1.5
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male examined closely has fewer set
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Figs. 26-29.-Texella kokoweej, new
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apical region which loses the apica
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Figs. 38-41.-Texella shoshone, new
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Distribution.-Known only from the t
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Figs. 52-55.-Texella brevistyla, ne
- Page 201 and 202: Texellajungi, new speciesFigs. 60-7
- Page 203 and 204: Figs. 66-69.-Texellajungi, new spec
- Page 205 and 206: than S; SA with laterobasal carina
- Page 207 and 208: ~81//J/'/ ;'?/ ~~.........--~~I, II
- Page 209 and 210: cylindrical, retina and cornea abse
- Page 211 and 212: Figs. 93-96.-Texella cokendolpheri,
- Page 213 and 214: SA with well developed prong and re
- Page 215 and 216: Figs. 105-108.-Texelia mulaiki Good
- Page 217 and 218: and McCarty Caves, which are known
- Page 219 and 220: Color orange. Body of medium rugosi
- Page 221 and 222: Notes.-The type locality was errone
- Page 223 and 224: Figs. 128-131.-Taella reyesi, new s
- Page 225 and 226: Figs. 136-139.-Texella reyesi, new
- Page 227 and 228: Figs. 144-147.-Texella reyesi, new
- Page 229 and 230: Figs. 152-155.-Texella reyesi, new
- Page 231 and 232: 1989 (W. Elliott, J. Reddell, and M
- Page 233 and 234: Table 3.-Continued.# locality sex S
- Page 235 and 236: mesoapical; patella, 2 mesal; tibia
- Page 237 and 238: Figs. 162-165.-Texella gmbbsi, new
- Page 239 and 240: Figs. 166-169.-Texella diplospina,
- Page 241 and 242: Figs. 174-177.-Texella renkesae, ma
- Page 243 and 244: Figs. 178-18\.-Teulla spinoperca, n
- Page 245 and 246: Distribution.-Known only from Fayet
- Page 247 and 248: Figs. 190-193.-Texellafendi, new sp
- Page 249 and 250: CLASSIFICAnONTexellabifurcata group
- Page 251: Chandler, D.S. 1992. The Pselaphida
- Page 255 and 256: stemite VI slightly impressed at ba
- Page 257 and 258: vertexal carinae, and the laterally
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- Page 261 and 262: small ventral carina near base, pro
- Page 263: Grigarick, A.A., and R.O. Schuster.
- Page 266 and 267: Cicurifla (Cicurella) holsiflgeri G