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Color yellowish orange. Body coarsely rugose;tubercles on eye mound, pars thoracica, tergitemargins, coxae, etc. Carapace with 2 pairs of AT.Eye mound low, rounded; eyes well developed.Palpal megaspines: trochanter, I ventral; femur, 1mesoapical; patella, 2 mesal; tibia, 2 mesal. Tarsalcount: 3-5-4-5. Tr IV spur relatively straight,length, 0.21. POP absent.Penis (Figs. 45-47): VPP apically pointed; with3 dorsal, 4? lateral, and 2 ventral setae; ASmedium-sized, stout. Glans: BK absent; MLapparently absent; PSL short, claw-like; PSL2vestigial, apically directed. S compressed; SAapparently represented by small, laterobasal carina;BF absent.Female: Unknown.Note.-The single specimen available for studywas unusually resistant in having its genitalia extractedand, unfortunately, was badly damaged inthe process.Natural History.-The holotype was collectedunder a rock (?granite) in a mesic southern chaparralcommunity along the canyon slope. A subsequentvisit to the locality, on 2 April 1985, failed to produceadditional specimens.Other Material Examined.-None.4344Figs. 42-47.-Texella species, male genitalia: 42-43, T. kokoweef, new species, paratopotype: 42, penis, suhventral view,expanded with KOH; 43, penis, lateral view. 44, T. shoshone, new species, paratopotype: Penis, lateral view, expanded with KOH.45-47, T. deserticola, new species, holotype: 45, penis, lateral view, expanded; 46, penis, lateral view, partially expanded; 47,penis, ventral view. Scale bar = 0.25 mm.186
Distribution.-Known only from the type locality(Map 1).The mulaiki GroupDiagnosis.-The following combination of malecharacters distinguishes the members of this group:POP present (secondarily lost in T. mulaiki,hardeni, bilobata, !eflai, and homi), glans peniswith BK (lost in T. brevistyla and the spiflopercainfragroup), and stylus with BF (small in T.brevistyla) (Figs. 7, 132, 133). No characters havebeen found which distinguish females.Range.-Known from southern New Mexico tocentral Texas (Maps 2-4).The brevistyla SubgroupDiagnosis.-The combination of S shorter thanPSL, and the absence of a SA (Figs. 52-55, 66-69,74-76) distinguishes males of these species from allother Texella.Range.-Known from western central Texas(Maps 3,4).Texella brevistyla, new speciesFigs. 48-59, 76.Diagnosis.-Males of this species are distinguishedfrom other Texella by the following combinationof characters: VPP apically convergent; PSLlong, attenuated, spiraled; S short, tube-like; BFpresent; SA absent; BK absent (Figs. 52-55, 76).Type.-Male holotype from Crom Cave, UvaldeCounty, Texas (Summer 1983; R. Waters), depositedin CAS.Etymology.- The speci fie name refers to theshort stylus characteristic of this species.Description.-Total body length, 1.56-2.18.Scute length, 1.18-1.44. Leg II length, 3.54-4.03.Leg II/Scute length, 2.58-3.17. (N = 11).Color orange to brownish orange. Body ofmedium rugosity; tubercles on eye mound, pars thoracica,tergite margins, and coxae. Carapace with 8pairs of AT arranged in two rows. Eye moundbroadly conical, eyes well developed, cervicalgroove prominent. Palpal megaspines: trochanter,1 ventral; femur, 1 mesoapical; patella, 2 mesal;tibia, 3 mesal. Tarsal count: 3-5-4-5.Male (holotype): Total body length, 1.85. Scutelength, 1.38; width, 1.44. Eye mound length, 0.41;width, 0.41. Leg II length, 3.56. TrIV spur robust,apically bent; length, 0.51. POP length, 0.26.Penis (Figs. 52-55, 76): VPP apically converging;with 2 dorsal, 10 lateral, and 4 ventral setae;AS short, curved. Glans: BK absent; ML present,of moderate sire; PSL elongate, tapering apically,spiraled. S short, tubular; BF present, reduced; SAabsent.Female (paratopotype): Total body length, 1.64.Scute length, 1.28; width, 1.33. Eye mound length,0.32; width, 0.31. Leg II length, 3.54. TrIV withtwo ventral tubercles.Ovipositor (Figs. 56, 58, 59): cuticle intricatelyfolded; dorsal surface sparsely set with microserrations,ventral smooth; 1 pair of small apical teethpresent; setal pattern: 1 pair dorsal, 4 pairs lateral, 1pair ventral.Variation.-The cervical groove is more prominentin specimens from Crom Cave. One male(SEM) has a short TrIV spur (L = 0.35) and astrongly reduced POP (L = 0.05) (Figs. 50, 51); inthe remaining males (6) these structures are uniformlylarge (about 0.50 and 0.30 respectively).The aberrant male is distinctly smaller than theothers (scute length = 1.18 compared to 1.31-1.44for the others). One male from BFS Cave (SEM)lacks an apical spine (Figs. 52-55) which, like thetip of one prong, may be broken. The ovipositor ofthe female from BFS Cave (SEM) lacks a pair ofapical teeth (Figs. 56, 58) which are present,although small, in all three females from CromCave.Note.-The PSL are spiral-shaped and enclosethe S in the unexpanded and partially expandedglans. With more complete expansion the PSL separateand become somewhat unspiraled (as in Fig.53).Natural History.-One female (Crom Cave) haswhat appears to be two large eggs (one visiblethrough the somewhat transparent sternites and thesecond revealed through dissection). The presumedeggs are quite large (0.36 x 0.46), much larger thanthe genital opening (0.18 x 0.23), and occupy muchof the abdominal cavity. We have seen specimens ofanother laniatorid harvestman, Hoplobunus species(Gonyleptoidea), from a nearby cave, Frio QueenCave.Other Material Examined (Paratypes).- UNITED STATES: Texas: Uvalde Co.: Crom Cave,Summer 1983 (R. Waters, TMM, CDU), 3 females;BFS Cave, 8 June 1989 (A. Grubbs, "A.c.", and R. Waters, TMM, CAS, CDU), 6 males, 1female.Distribution.-Known only from Crom and BFScaves, Uvalde County, Texas (Maps 2-4).187
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PREFACEThe present volume is the se
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TABLE OF CONTENTSHOLSINGER, JOHN R.
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the state of Coahuila in northern M
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Fig. 2.-Anesia welboumi, new specie
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\\. \ - -' ..........---~\ I// --..
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Mexiweckelia hardeni, new speciesFi
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2 sets of 1 or 2 setae each; dactyl
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AFig. 7.-Holsingerius smaragdinus,
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have as many setae on the inner pla
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Antenna 1 about 33 % length of body
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Of biogeographic interest for the h
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Bowman, T .E. 1992. Two subterranea
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A note by Scott Harden that accompa
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~~..~ ~~ I))\\ "-,'.=bFig. 3.-Speoc
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unarmed, except in pleopod 2, which
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Cokendolpher, LC., and l.R. Reddell
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lobes. The larger setae vary greatl
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the relationships of the order and
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zomids their absence is considered
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some species could be either split,
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have small pores over the surface o
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inhabits tropical deciduous forest
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huitvnolotitlensis from A. stygius
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8-10, figs. 5-7; Rowland, 1973c:136
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(in row) and one pair large posteri
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(0.34); tarsus 0.64 (0.58); total 4
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Abdomen: Tergite I with two pairs a
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setae near posterior margin. stemit
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setae, and ten ventral setae. Stemi
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1,980 m elev., 26 Dec. 1986 (T. Tre
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asitarsal-tarsal proportions: 15:4:
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and one pair setae at base of proce
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Cephalothorax: Propeltidium 1.66 mm
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Male adult unknown.Immature paratyp
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IV: trochanter 1.20 (1.10); femur 3
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Cokendolpher, 1.C. 1981. The order
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Gertsch, W.J. 1992. Distribution pa
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same families and genera, but also
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species from the United States and
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the posterior median pair short and
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great many species of North America
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Key to the Eyed Females1. Eight eye
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39111012Figs. 1-12.-Ventral and dor
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1419 23Figs. 13-24.-Ventral and dor
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27 293334Figs. 25-36.-Ventral and d
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Figs. 37-48.-Ventral and retrolater
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Cicurina blanco, new speciesFigs. 7
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Type-data.-Female holotype from ins
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Description.-Female holotype: Lengt
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lengths: first femur 2 rom, fourth
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Cicurina pablo, new speciesFigs. 10
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canal in nearly vertical posItion;
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Cicurina vespera, new speciesFigs.
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procurved canal across sac; connect
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Cicurina caverna, new speciesFigs.
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with sac of similar size set in obl
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February 1964 (J. Reddell, D. McKen
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County: Diamond Cave, 16 August 196
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Roth, V.D. 1992. A new and first tr
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LITERATURE CITEDBarr, T.C. 1963. Ec
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Muchmore, W.B. 1992. Cavernicolous
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Species of Aphrastochthonius are kn
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FAMILY NEOBISIIDAE CHAMBERLINGenus
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trochanter 2.5 (2.6), femur 5.55 (5
Page 145 and 146: ottom of entrance pit, Ogle Cave (O
Page 147 and 148: tactile seta on tibia and basitarsu
Page 149 and 150: Missouri may be conspecific (unpubl
Page 151 and 152: Trichobothriotaxy of chela generall
Page 153 and 154: Female (figures given first for all
Page 155 and 156: transverse furrows; eyespots not ev
Page 157 and 158: Reddell and W. Russell); I female f
Page 159 and 160: and Acuminochernes, along with the
Page 161 and 162: Chamberlin, J.C. 1946. The genera a
Page 163: (0.36); chela (without pedicel) 2.0
Page 167 and 168: although not recently studied is no
Page 169 and 170: and stylar outgrowths, present in s
Page 171 and 172: in Phalangodes (et al.) is thick an
Page 173 and 174: pairs) are found in six species: T.
Page 175 and 176: elated species IS probably best int
Page 177 and 178: legs than expected (2.6-3.2). This
Page 179 and 180: ~ 3.3 are synapomorphic); all trogl
Page 181 and 182: TAXONOMYTEXELLA Goodnight and Goodn
Page 183 and 184: 14. BK absent (Figs. 177, 180). SA
Page 185 and 186: Figs. 8-11.-Texella bijUrcata (Brig
Page 187 and 188: Description.-Total body length, 1.5
Page 189 and 190: male examined closely has fewer set
Page 191 and 192: Figs. 26-29.-Texella kokoweej, new
Page 193 and 194: apical region which loses the apica
Page 195: Figs. 38-41.-Texella shoshone, new
Page 199 and 200: Figs. 52-55.-Texella brevistyla, ne
Page 201 and 202: Texellajungi, new speciesFigs. 60-7
Page 203 and 204: Figs. 66-69.-Texellajungi, new spec
Page 205 and 206: than S; SA with laterobasal carina
Page 207 and 208: ~81//J/'/ ;'?/ ~~.........--~~I, II
Page 209 and 210: cylindrical, retina and cornea abse
Page 211 and 212: Figs. 93-96.-Texella cokendolpheri,
Page 213 and 214: SA with well developed prong and re
Page 215 and 216: Figs. 105-108.-Texelia mulaiki Good
Page 217 and 218: and McCarty Caves, which are known
Page 219 and 220: Color orange. Body of medium rugosi
Page 221 and 222: Notes.-The type locality was errone
Page 223 and 224: Figs. 128-131.-Taella reyesi, new s
Page 225 and 226: Figs. 136-139.-Texella reyesi, new
Page 231 and 232: 1989 (W. Elliott, J. Reddell, and M
Page 233 and 234: Table 3.-Continued.# locality sex S
Page 235 and 236: mesoapical; patella, 2 mesal; tibia
Page 237 and 238: Figs. 162-165.-Texella gmbbsi, new
Page 239 and 240: Figs. 166-169.-Texella diplospina,
Page 241 and 242: Figs. 174-177.-Texella renkesae, ma
Page 243 and 244: Figs. 178-18\.-Teulla spinoperca, n
Page 245 and 246: Distribution.-Known only from Fayet
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Figs. 190-193.-Texellafendi, new sp
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CLASSIFICAnONTexellabifurcata group
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Chandler, D.S. 1992. The Pselaphida
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Key to Species1. Abdominal segments
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stemite VI slightly impressed at ba
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vertexal carinae, and the laterally
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is associated with rotten woods (Ch
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small ventral carina near base, pro
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Grigarick, A.A., and R.O. Schuster.
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Cicurifla (Cicurella) holsiflgeri G
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