Description.-Length 2.20. Males with antennaltubercles on head close, <strong>for</strong>ming a broad tubercleconstricted at base, vertexal foveae a distance of oneocular facet from eyes, eyes with about eight largefacets, maxillary palpi with last three segments elongate,together nearly as long as antennae, penultimatesegment angulate near base on mesal margin.Pronotum lacking any sulci, with small medianand lateral antebasal foveae. Elytra with two basalfoveae, lacking subhumeral fovea. Metastemumwith short longitudinal carina extending anteriorlyfrom near inner margins of each metacoxa; stemitesII-VI broadly flattened at middle; protrochanterswith prominent spine, metatrochanters with elongatebroad flange extending ventrally.Females are unknown.Male specimen from Gorman <strong>Cave</strong>, Texas: antenna1.19, metafemur 0.76, metatibia 0.79, metatarsus,0.47.Specimen examined.-Texas: San Saba County,Gorman <strong>Cave</strong>, 6 mi. SE Bend on Colorado River,III-15-1963, J. R. Reddell (DSC).Comments.-Cylindrarctus bicornis was takenfrom washed-in organic debris in total darkness severalhundred meters from the cave entrance (JamesR. Reddell, pers. carom.). This is the second knownspecimen of this species. The other specimen, theholotype, was produced by Belfrage from "Texas,"where he had collected most extensively at Cliftonand Norse in Bosque County. The other members ofthe genus are most commonly encountered in moistlitter near water, and bicornis will probably only befound in caves at their entrances since it lacks anyobvious cave adapted features. Cylindrarctus bicornisis readily separated from the other cave pselaphidsby the elongate last three segments of themaxillary palpi that together are nearly equal to theantennae in length.10. Tmesiphorus costalis LeConte(Fig. 21)Tmesiphorus costalis LeConte, 1849:77. Type locality:Pennsylvania. Holotype male (MCZC).Description.-Length 3.08-3.52. Males withhead and pronotum coarsely punctate, antennomereIX cylindrical and three times length of VIII, withlongitudinal groove on outer face, X slightly shorterthan IX and flattened ventrally with anterodistal angletuberculate, XI excavate in basal third on ventralsurface; maxillary palpi with apical segment triangular,previous two segments with lateral spine.Pronotal disc rounded; elytra with two basal foveae,lacking subhumeral fovea; abdomen with sternitesII-III lightly flattened at middle.Females with apical antennomeres narrower,lacking modifications; abdominal sternites broadlyrounded.Male specimen from Puberty Pit, Texas: antennae1.68, metafemora 1.04, metatibiae 1.20, metatarsi0.48.Specimens examined.-TEXAS: Burnet County:Simon Says Sink, XI-12-1990, J. Reddell, M.Reyes, berlese litter (2, TMMC); Snake Pit Sink,XI-20-1990, J. Reddell, M. Reyes, underside ofrock loosely buried in silt (2, DSC); SimonsSquirm-Around <strong>Cave</strong>, XI-20-1990, J. Reddell, M.Reyes, underside of rocks loosely buried in silt (4,DSC); Shin Oak Sink, 1-17-1991, J. Reddell, M.Reyes (4" DSC). San Saba County: Puberty Pit,IV-21-1990, D. Allen, W. Elliott, B. Fralia (1,TMMC). Travis County: Big Oak <strong>Cave</strong>, X-17-1990,J. Reddell, M. Reyes, on underside of rock indarkness (I, TMMC); Twin Dig Pit, XII-1l-1990,J. Reddell, berlese rodent nests at lowest point ofcave in total darkness (1, TMMC).Comments.-This species has been collected invarious leaf litters, under bark, with the antAphaenogaster fulva Roger (Park, 1933), and onespecimen has been found in a cave (Park, 1960).Tmesiphorus costalis and the other Nearctic speciesin the genus, carinatus (Say), are easily recognizedby the <strong>for</strong>m of the maxillary palpi, and the coarselypunctate head and pronotum. Tmesiphorus costalis islarger than carinatus, the latter being easily distinguishedfrom costalis by a median longitudinalcarina on the first visible abdominal tergite.11. Hamotus (Hamotoides) electrae Park(Figs. 22-23)Hamotus electrae Park, 1942:327. Type locality:San Juan, Veracruz, MEXICO. Holotype male(FMNH).Description.-Length 3.20-3.40. Males with antennomeresIV-VIII transverse, IX twice as long asVIII, IX slightly shorter than X, both IX and Xslightly longer than wide; last segment of maxillarypalpus enlarged, twice as long as wide, with longitudinalsulcus on mesal margin.Pronotum with distinct transverse antebasal sulcus,width about half that of median antebasal fovea;elytra with two basal foveae, lacking subhumeral fovea;protrochanters carinate ventrally, a few specimenswith small denticle on carina, profemora with250
small ventral carina near base, protibiae with smalltooth on mesal margin at point about two-thirds oflength, mesotrochanters vaguely carinate on ventralmargin, metasternum deeply and transversely impressedin apical two-thirds, posterior margin raisedas flanges at mesal margins of metacoxae; abdominalsternites II-VI widely impressed.Females lack modifications of legs and metasternum;abdominal sternites broadly rounded.Male specimen from Porcupine <strong>Cave</strong>, Texas: antennae1.44, metafemora 1.00, metatibiae 1.08,metatarsi 0.54.Specimens examined.-3: TEXAS: KinneyCounty, Porcupine <strong>Cave</strong>, X-17-1987, G. Veni, J.Ivy (DSC and TMMC). Five other specimens werecollected at Bentsen-Rio Grande St. Pk., VI-I01975, by Robert Turnbow at ultraviolet light (DSCand RHT).Comments.-This species was previously knownonly from the holotype male collected in Veracruz,Mexico (Park, 1942). It is apparently a free-livingspecies, and will probably only be found when associatedwith caves at their entrances. Park (1942)placed this species in Group XII of Hamotus, wherethe other four North American species are placed.The males of electrae are readily distinguished fromall other Hamotus (Hamotoides) species by thetransverse impression of the metasternum, and themetasternal laminae projecting posteriorly near themesal margins of the metacoxae in males.Trimioiircus musamator n. sp.(Figs. 24-26)Description.-Length 1.16-1.24. Male head withvertex extended and flattened laterally, eyes not~~ Of;;18oClI/, r '22Figs. 18-26.-18-19, Texamaurops reMelli: 18, posterior view, right metatibia of female; 19, dorsal and right lateral view,aedeagus. 20, Cylindrarctus bicomis, dorsal view, male head. 21, Tmesiphorns costalis, dorsal view, male head. 22-23, Hamotuselectrae: 22, dorsal view, male head; 23, dorsal and right lateral view, aedeagus. 24-26, Trimioarcus musamator: 24, dorsal view,male head; 25 posterior view, male right middle leg; 26, dorsal and right lateral view, aedeagus. Scale line equals 0.1.251
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PREFACEThe present volume is the se
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TABLE OF CONTENTSHOLSINGER, JOHN R.
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the state of Coahuila in northern M
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Fig. 2.-Anesia welboumi, new specie
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\\. \ - -' ..........---~\ I// --..
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Mexiweckelia hardeni, new speciesFi
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2 sets of 1 or 2 setae each; dactyl
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AFig. 7.-Holsingerius smaragdinus,
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have as many setae on the inner pla
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Antenna 1 about 33 % length of body
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Of biogeographic interest for the h
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Bowman, T .E. 1992. Two subterranea
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A note by Scott Harden that accompa
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~~..~ ~~ I))\\ "-,'.=bFig. 3.-Speoc
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unarmed, except in pleopod 2, which
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Cokendolpher, LC., and l.R. Reddell
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lobes. The larger setae vary greatl
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the relationships of the order and
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zomids their absence is considered
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some species could be either split,
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have small pores over the surface o
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inhabits tropical deciduous forest
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huitvnolotitlensis from A. stygius
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8-10, figs. 5-7; Rowland, 1973c:136
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(in row) and one pair large posteri
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(0.34); tarsus 0.64 (0.58); total 4
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Abdomen: Tergite I with two pairs a
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setae near posterior margin. stemit
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setae, and ten ventral setae. Stemi
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1,980 m elev., 26 Dec. 1986 (T. Tre
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asitarsal-tarsal proportions: 15:4:
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and one pair setae at base of proce
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Cephalothorax: Propeltidium 1.66 mm
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Male adult unknown.Immature paratyp
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IV: trochanter 1.20 (1.10); femur 3
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Cokendolpher, 1.C. 1981. The order
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Gertsch, W.J. 1992. Distribution pa
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same families and genera, but also
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species from the United States and
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the posterior median pair short and
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great many species of North America
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Key to the Eyed Females1. Eight eye
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39111012Figs. 1-12.-Ventral and dor
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1419 23Figs. 13-24.-Ventral and dor
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27 293334Figs. 25-36.-Ventral and d
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Figs. 37-48.-Ventral and retrolater
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Cicurina blanco, new speciesFigs. 7
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Type-data.-Female holotype from ins
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Description.-Female holotype: Lengt
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Description.-Female holotype: Lengt
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lengths: first femur 2 rom, fourth
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Cicurina pablo, new speciesFigs. 10
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Description.-Female holotype: Lengt
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canal in nearly vertical posItion;
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Cicurina vespera, new speciesFigs.
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procurved canal across sac; connect
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Cicurina caverna, new speciesFigs.
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with sac of similar size set in obl
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February 1964 (J. Reddell, D. McKen
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County: Diamond Cave, 16 August 196
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Roth, V.D. 1992. A new and first tr
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LITERATURE CITEDBarr, T.C. 1963. Ec
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Muchmore, W.B. 1992. Cavernicolous
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Species of Aphrastochthonius are kn
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FAMILY NEOBISIIDAE CHAMBERLINGenus
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trochanter 2.5 (2.6), femur 5.55 (5
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ottom of entrance pit, Ogle Cave (O
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tactile seta on tibia and basitarsu
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Missouri may be conspecific (unpubl
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Trichobothriotaxy of chela generall
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Female (figures given first for all
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transverse furrows; eyespots not ev
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Reddell and W. Russell); I female f
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and Acuminochernes, along with the
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Chamberlin, J.C. 1946. The genera a
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(0.36); chela (without pedicel) 2.0
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although not recently studied is no
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and stylar outgrowths, present in s
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in Phalangodes (et al.) is thick an
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pairs) are found in six species: T.
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elated species IS probably best int
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legs than expected (2.6-3.2). This
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~ 3.3 are synapomorphic); all trogl
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TAXONOMYTEXELLA Goodnight and Goodn
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14. BK absent (Figs. 177, 180). SA
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Figs. 8-11.-Texella bijUrcata (Brig
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Description.-Total body length, 1.5
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male examined closely has fewer set
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Figs. 26-29.-Texella kokoweej, new
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apical region which loses the apica
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Figs. 38-41.-Texella shoshone, new
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Distribution.-Known only from the t
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Figs. 52-55.-Texella brevistyla, ne
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Texellajungi, new speciesFigs. 60-7
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Figs. 66-69.-Texellajungi, new spec
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than S; SA with laterobasal carina
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~81//J/'/ ;'?/ ~~.........--~~I, II
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- Page 211 and 212: Figs. 93-96.-Texella cokendolpheri,
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- Page 215 and 216: Figs. 105-108.-Texelia mulaiki Good
- Page 217 and 218: and McCarty Caves, which are known
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- Page 221 and 222: Notes.-The type locality was errone
- Page 223 and 224: Figs. 128-131.-Taella reyesi, new s
- Page 225 and 226: Figs. 136-139.-Texella reyesi, new
- Page 227 and 228: Figs. 144-147.-Texella reyesi, new
- Page 229 and 230: Figs. 152-155.-Texella reyesi, new
- Page 231 and 232: 1989 (W. Elliott, J. Reddell, and M
- Page 233 and 234: Table 3.-Continued.# locality sex S
- Page 235 and 236: mesoapical; patella, 2 mesal; tibia
- Page 237 and 238: Figs. 162-165.-Texella gmbbsi, new
- Page 239 and 240: Figs. 166-169.-Texella diplospina,
- Page 241 and 242: Figs. 174-177.-Texella renkesae, ma
- Page 243 and 244: Figs. 178-18\.-Teulla spinoperca, n
- Page 245 and 246: Distribution.-Known only from Fayet
- Page 247 and 248: Figs. 190-193.-Texellafendi, new sp
- Page 249 and 250: CLASSIFICAnONTexellabifurcata group
- Page 251 and 252: Chandler, D.S. 1992. The Pselaphida
- Page 253 and 254: Key to Species1. Abdominal segments
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- Page 259: is associated with rotten woods (Ch
- Page 263: Grigarick, A.A., and R.O. Schuster.
- Page 266 and 267: Cicurifla (Cicurella) holsiflgeri G