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searchable PDF - Association for Mexican Cave Studies

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indicating facet remnants; antennomere X with largefovea covering ventral half.Pronotum with median longitudinal sulcus faintin basal half of disc, disc sparsely and minutelypunctate; elytra with 3 basal foveae and subhumeralfovea; stemite VI with lateral setate tubercles atapex, disc between tubercles flat; second mesotarsomeresstraight.Females lacking modifications of head, antennae,and sixth stemite; frons and anterior half of vertexcoarsely punctate, posterior half of vertex smoothlyroughened.Holotype male: antennae 1.40, metafemora 0.96,metatibiae 1.00, metatarsi 0.48.Specimens examined.-6: TEXAS: HOLO­TYPE male, Hays County, Grapevine <strong>Cave</strong>,VII-I-1990, A. Grubbs, J. Reddell, M. Reyes(FMNH). PARATYPES, all same locality: 1 female,eutopotypical (TMMC); 2 males, 1 female,VI-IO-1990, A. Grubbs, L. Davis, J. Elliott (DSC,TMMC); 1 female, VII-9-1990, A. G. Grubbs(DSC). The specimens collected on June 10 areteneral.Etymology.-The species is named <strong>for</strong> theprincipal collector of the series, Andrew Grubbs.Comments.-The type series was collected underwashed-in leaf litter at the litter-clay floor interfacefrom the terminal room of Grapevine <strong>Cave</strong> in totaldarkness (James R. Reddell, pers. comm.). Themale modifications of the head are generally similarto those of clypeonotus, but differ in the <strong>for</strong>m of thevertexal apex, lack of lateral clypeal spines, and inthe <strong>for</strong>m of antennomeres I and X. The male antennae,median clypeal spine, and <strong>for</strong>m of the aedeagusare also similar to those of venyivi, but may be readilyseparated by the truncate vertexal apex andcoarse punctation on the anterior half of the vertex.4. Batrisodes (Excavodes) reyesi, n. sp.(Figs. 7-8)Description.-Length 2.36-2.44. Males withhead transversely excavate just anterior to antennalbases, frontal hom prominent and with dense shortsetae directed posterolaterally, anterior margin ofclypeus broadly rounded, anterior half of vertexwith obscure transverse rugules and smooth onposterior half, anterior margin of vertex weaklybiemarginate over transverse excavation, short setaealong margin longest near lateral edges, stronglateral carinae of vertex extending anteriorly toantennal bases, median carina extending from baseto just anterior to line between nude vertexal foveae,eyes represented by a number of small scatteredfacet rudiments appearing as granules; first antennomere(scape) angulate on ventral margin, denselyand minutely punctate in lower half, antennomere Xwith large fovea occupying over half of ventralsurface.Pronotum smooth, with median longitudinal sulcusextending to near pronotal apex; elytra withthree basal foveae and subhumeral fovea; metasternumat middle with small deep circular impressionjust anterior to metacoxae; stemite VI medially impressedwith tuberculate raised areas laterally, lineof thickened setae extending from tubercles tomiddle; second mesotarsomeres straight.Females lack modifications of head, antennae,and stemite VI; vertex depressed between antennalbases and smoothly rugulose; metastemal impressionpresent.Male holotype: antennae 1.52, metafemora 1.04,metatibiae 1.05, metatarsi 0.44.Specimens examined.-13: HOLOTYPE male,Texas, Burnet County, Fenceline Sink, IV-17-1990,J. Reddell, M. Reyes (FMNH). PARATYPES: 1female, eutopotypical (TMMC); I male, same locality,V-27-1989, M. Reyes (DSC); 1 male, Snake PitSink, XI-20-1990, J. Reddell, M. Reyes, undersideof rock loosely buried in silt (TMMC). TravisCounty: 1 female, Armadillo Ranch Sink,IX-23-1990, J. Reddell, M. Reyes, C. Sexton, fromcrack in rotten wood (DSC); 3 females, YellowBerry <strong>Cave</strong>, XII-I 11990, J. Reddell, M. Reyes(DSC, TMMC); 2 males, 3 females, Moss Pit,III-5-1991, J. Reddell, M. Reyes, underside ofrocks deeply buried in silt (DSC, TMMC). The twospecimens collected in April are teneral.Etymology.-The name is derived from MarcelinoReyes, the principal collector of the type series.Comments.-Batrisodes reyesi was found on theunderside of rocks deeply buried in silt at thebottom of the entrance drop of Fenceline Sink,associated with troglobitic spiders, Cicurina (Cicurella)species, and millipedes, Speodesmus species;in Yellow Berry <strong>Cave</strong> beneath rocks at the bottom ofthe second drop in total darkness; and in Moss Piton the underside of rocks deeply buried in silt atcave bottom in association with blind isopods of thefamily Trichoniscidae (James R. Reddell, pers.comm.).The general <strong>for</strong>m of the modified vertex and firstantennomeres are similar to those of clypeo/lotus(Brendel), a myrmecophile with reduced eyes. Themales of reyesi are easily distinguished by the relativelyflat vertex, the stronger lateral and median246

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