indicating facet remnants; antennomere X with largefovea covering ventral half.Pronotum with median longitudinal sulcus faintin basal half of disc, disc sparsely and minutelypunctate; elytra with 3 basal foveae and subhumeralfovea; stemite VI with lateral setate tubercles atapex, disc between tubercles flat; second mesotarsomeresstraight.Females lacking modifications of head, antennae,and sixth stemite; frons and anterior half of vertexcoarsely punctate, posterior half of vertex smoothlyroughened.Holotype male: antennae 1.40, metafemora 0.96,metatibiae 1.00, metatarsi 0.48.Specimens examined.-6: TEXAS: HOLOTYPE male, Hays County, Grapevine <strong>Cave</strong>,VII-I-1990, A. Grubbs, J. Reddell, M. Reyes(FMNH). PARATYPES, all same locality: 1 female,eutopotypical (TMMC); 2 males, 1 female,VI-IO-1990, A. Grubbs, L. Davis, J. Elliott (DSC,TMMC); 1 female, VII-9-1990, A. G. Grubbs(DSC). The specimens collected on June 10 areteneral.Etymology.-The species is named <strong>for</strong> theprincipal collector of the series, Andrew Grubbs.Comments.-The type series was collected underwashed-in leaf litter at the litter-clay floor interfacefrom the terminal room of Grapevine <strong>Cave</strong> in totaldarkness (James R. Reddell, pers. comm.). Themale modifications of the head are generally similarto those of clypeonotus, but differ in the <strong>for</strong>m of thevertexal apex, lack of lateral clypeal spines, and inthe <strong>for</strong>m of antennomeres I and X. The male antennae,median clypeal spine, and <strong>for</strong>m of the aedeagusare also similar to those of venyivi, but may be readilyseparated by the truncate vertexal apex andcoarse punctation on the anterior half of the vertex.4. Batrisodes (Excavodes) reyesi, n. sp.(Figs. 7-8)Description.-Length 2.36-2.44. Males withhead transversely excavate just anterior to antennalbases, frontal hom prominent and with dense shortsetae directed posterolaterally, anterior margin ofclypeus broadly rounded, anterior half of vertexwith obscure transverse rugules and smooth onposterior half, anterior margin of vertex weaklybiemarginate over transverse excavation, short setaealong margin longest near lateral edges, stronglateral carinae of vertex extending anteriorly toantennal bases, median carina extending from baseto just anterior to line between nude vertexal foveae,eyes represented by a number of small scatteredfacet rudiments appearing as granules; first antennomere(scape) angulate on ventral margin, denselyand minutely punctate in lower half, antennomere Xwith large fovea occupying over half of ventralsurface.Pronotum smooth, with median longitudinal sulcusextending to near pronotal apex; elytra withthree basal foveae and subhumeral fovea; metasternumat middle with small deep circular impressionjust anterior to metacoxae; stemite VI medially impressedwith tuberculate raised areas laterally, lineof thickened setae extending from tubercles tomiddle; second mesotarsomeres straight.Females lack modifications of head, antennae,and stemite VI; vertex depressed between antennalbases and smoothly rugulose; metastemal impressionpresent.Male holotype: antennae 1.52, metafemora 1.04,metatibiae 1.05, metatarsi 0.44.Specimens examined.-13: HOLOTYPE male,Texas, Burnet County, Fenceline Sink, IV-17-1990,J. Reddell, M. Reyes (FMNH). PARATYPES: 1female, eutopotypical (TMMC); I male, same locality,V-27-1989, M. Reyes (DSC); 1 male, Snake PitSink, XI-20-1990, J. Reddell, M. Reyes, undersideof rock loosely buried in silt (TMMC). TravisCounty: 1 female, Armadillo Ranch Sink,IX-23-1990, J. Reddell, M. Reyes, C. Sexton, fromcrack in rotten wood (DSC); 3 females, YellowBerry <strong>Cave</strong>, XII-I 11990, J. Reddell, M. Reyes(DSC, TMMC); 2 males, 3 females, Moss Pit,III-5-1991, J. Reddell, M. Reyes, underside ofrocks deeply buried in silt (DSC, TMMC). The twospecimens collected in April are teneral.Etymology.-The name is derived from MarcelinoReyes, the principal collector of the type series.Comments.-Batrisodes reyesi was found on theunderside of rocks deeply buried in silt at thebottom of the entrance drop of Fenceline Sink,associated with troglobitic spiders, Cicurina (Cicurella)species, and millipedes, Speodesmus species;in Yellow Berry <strong>Cave</strong> beneath rocks at the bottom ofthe second drop in total darkness; and in Moss Piton the underside of rocks deeply buried in silt atcave bottom in association with blind isopods of thefamily Trichoniscidae (James R. Reddell, pers.comm.).The general <strong>for</strong>m of the modified vertex and firstantennomeres are similar to those of clypeo/lotus(Brendel), a myrmecophile with reduced eyes. Themales of reyesi are easily distinguished by the relativelyflat vertex, the stronger lateral and median246
vertexal carinae, and the laterally tuberculate stemiteVI.5. Batrisodes (Excavodes) texanus n. sp.(Figs. 9-12)Description.-Length 2.60-2.88. Male head withvague transverse impression anterior to antennalbases, impression shallow and medianly angulate,vertex smooth and sparsely setate, antennal tuberclesprominent with a few coarse punctures dorsally,sides of head smoothly curved and flat with a fewgranules present where eyes should be, lateral carinaeextending sinuately from head base of outer angieof antennal tubercles, median carina extendinganteriorly to point between nude vertexal foveae; antennomeresall elongate, X nearly twice as wide asIX and narrowing in apical half, with large nude foveacovering one-third of surface in ventral view.Pronotum with median longitudinal sulcus shallowon disc; elytra with three basal foveae and subhumeralfovea; second mesotarsomeres straight.Females lacking transverse sulcus anterior to antennalbases, vertex merging smoothly with frons,antennomere X barely wider and longer than IX.Male holotype: antenna 1.63, metafemur, 1.32,mesofemur 1.32, metatarsi lost.Specimens examined.-4: TEXAS: WilliamsonCounty: HOLOTYPE male, Inner Space <strong>Cave</strong>rns(=Laubach's <strong>Cave</strong>), V-23-1965, W. Russell(FMNH). PARATYPES (all females): same locality,VIII-1968, W. Elliott (DSC); Off Campus<strong>Cave</strong>, IV-8- 1989, W. Elliott, J. Reddell, M. Reyes(DSC); Coffin <strong>Cave</strong>, 10 mi. NW Georgetown,XI-3-1963, J. Reddell (TMMC).Etymology.-The name is derived from itsknown occurrence restricted to Texas.Comments.-Batrisodes texanus was found inOff Campus <strong>Cave</strong> on the underside of a rock lightlyburied in silty clay in total darkness (James R. Reddell,pers. comm.). Since this species was includedas the Coffm <strong>Cave</strong> population of Texamaurops reddelliwhen reddelli was listed as an endangered species(Chambers and Jahrsdoerfer, 1988), texanusshould also be considered endangered under federallaw although no specific ruling has yet been published(Steve Chambers, pers. comm. to James R.Reddell).This species possesses elongate legs andantennae, and is superficially similar to T. reddelli.The lack of any ocular projection, and the presenceof the pencil of setae at the apex of the metatibiaereadily separate the two taxa. The aedeagus of thisspecies was figured by Barr (1974) as T. reddelli.Batrisodes texanus is placed in the subgenus Excavodesdue to the modified head anterior to theantennae and the straight mesotarsomeres of themale.6. Batrisodes (Excavodes) venyivi n. sp.(Figs. 13-14)Description.-Length 2.24. Males with headtransversely excavate just anterior to antennal bases,frontal hom prominent with short dense setae directedposterolaterally, anterior margin of clypeusangulate, anterior margin of vertex rounded withshallow medial emargination, margin anterior to antennalbases depressed, vertex smooth and lackinglateral and medial basal carinae; first antennomere(scape) not modified, antennomere X with fovea occupyingone-fourth of face in ventral view.Pronotum smooth, with median longitudinal sulcusbarely attaining middle of disk; elytra with threebasal foveae, lacking subhumeral fovea; stemite VIlight!Y impressed medially, smoothly granulate inimpression; second mesotarsomeres straight.Female unknown.Male holotype: antennae 1.05, metafemora 0.76,metatibiae 0.80, metatarsi 0.44.Specimen examined.-TEXAS: Bexar County:HOLOTYPE male, Helotes Hilltop <strong>Cave</strong>, IX-291984, J. Ivy, G. Veni (FMNH).Etymology.-The name is <strong>for</strong>med by combinationof the names of the collectors.Comments.-It is not close to any other speciesof Excavodes that I have seen. It is easily distinguishedby the lack of vertexal carinae, smooth vertex,and lack of a subhumeral fovea on the elytra.7. Batrisodes (Excavodes) globosus (LeConte)(Figs. 15, 16)Batrisus globosus LeConte, 1849: 100. Type localities:Pennsylvania and Georgia. Syntypes maleand female (MCZC).Description.-Length 2.36-2.48. Males withhead transversely excavated anterior to antennalbases, frons with acute medial tubercle bearing foursmall recurved setae at apex and longer laterally directedsinuate setae toward base, frontal tuberclenearly obscured in dorsal view by broadly roundedprojecting vertexal lobe, ventral margin of vertexallobe with small group of setae projecting ventrallylateral to frontal tubercle, vertex coarsely punctatein anterior third, generally smooth in posteriorportion with setae arising from small granules,247
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PREFACEThe present volume is the se
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TABLE OF CONTENTSHOLSINGER, JOHN R.
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the state of Coahuila in northern M
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Fig. 2.-Anesia welboumi, new specie
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\\. \ - -' ..........---~\ I// --..
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Mexiweckelia hardeni, new speciesFi
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2 sets of 1 or 2 setae each; dactyl
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AFig. 7.-Holsingerius smaragdinus,
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have as many setae on the inner pla
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Antenna 1 about 33 % length of body
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Of biogeographic interest for the h
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Bowman, T .E. 1992. Two subterranea
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A note by Scott Harden that accompa
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~~..~ ~~ I))\\ "-,'.=bFig. 3.-Speoc
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unarmed, except in pleopod 2, which
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Cokendolpher, LC., and l.R. Reddell
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lobes. The larger setae vary greatl
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the relationships of the order and
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zomids their absence is considered
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some species could be either split,
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have small pores over the surface o
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inhabits tropical deciduous forest
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huitvnolotitlensis from A. stygius
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8-10, figs. 5-7; Rowland, 1973c:136
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(in row) and one pair large posteri
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(0.34); tarsus 0.64 (0.58); total 4
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Abdomen: Tergite I with two pairs a
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setae near posterior margin. stemit
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setae, and ten ventral setae. Stemi
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1,980 m elev., 26 Dec. 1986 (T. Tre
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asitarsal-tarsal proportions: 15:4:
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and one pair setae at base of proce
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Cephalothorax: Propeltidium 1.66 mm
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Male adult unknown.Immature paratyp
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IV: trochanter 1.20 (1.10); femur 3
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Cokendolpher, 1.C. 1981. The order
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Gertsch, W.J. 1992. Distribution pa
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same families and genera, but also
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species from the United States and
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the posterior median pair short and
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great many species of North America
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Key to the Eyed Females1. Eight eye
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39111012Figs. 1-12.-Ventral and dor
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1419 23Figs. 13-24.-Ventral and dor
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27 293334Figs. 25-36.-Ventral and d
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Figs. 37-48.-Ventral and retrolater
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Cicurina blanco, new speciesFigs. 7
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Type-data.-Female holotype from ins
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Description.-Female holotype: Lengt
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Description.-Female holotype: Lengt
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lengths: first femur 2 rom, fourth
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Cicurina pablo, new speciesFigs. 10
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Description.-Female holotype: Lengt
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canal in nearly vertical posItion;
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Cicurina vespera, new speciesFigs.
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procurved canal across sac; connect
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Cicurina caverna, new speciesFigs.
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with sac of similar size set in obl
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February 1964 (J. Reddell, D. McKen
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County: Diamond Cave, 16 August 196
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Roth, V.D. 1992. A new and first tr
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LITERATURE CITEDBarr, T.C. 1963. Ec
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Muchmore, W.B. 1992. Cavernicolous
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Species of Aphrastochthonius are kn
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FAMILY NEOBISIIDAE CHAMBERLINGenus
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trochanter 2.5 (2.6), femur 5.55 (5
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ottom of entrance pit, Ogle Cave (O
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tactile seta on tibia and basitarsu
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Missouri may be conspecific (unpubl
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Trichobothriotaxy of chela generall
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Female (figures given first for all
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transverse furrows; eyespots not ev
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Reddell and W. Russell); I female f
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and Acuminochernes, along with the
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Chamberlin, J.C. 1946. The genera a
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(0.36); chela (without pedicel) 2.0
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although not recently studied is no
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and stylar outgrowths, present in s
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in Phalangodes (et al.) is thick an
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pairs) are found in six species: T.
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elated species IS probably best int
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legs than expected (2.6-3.2). This
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~ 3.3 are synapomorphic); all trogl
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TAXONOMYTEXELLA Goodnight and Goodn
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14. BK absent (Figs. 177, 180). SA
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Figs. 8-11.-Texella bijUrcata (Brig
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Description.-Total body length, 1.5
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male examined closely has fewer set
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Figs. 26-29.-Texella kokoweej, new
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apical region which loses the apica
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Figs. 38-41.-Texella shoshone, new
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Distribution.-Known only from the t
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Figs. 52-55.-Texella brevistyla, ne
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Texellajungi, new speciesFigs. 60-7
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Figs. 66-69.-Texellajungi, new spec
- Page 205 and 206: than S; SA with laterobasal carina
- Page 207 and 208: ~81//J/'/ ;'?/ ~~.........--~~I, II
- Page 209 and 210: cylindrical, retina and cornea abse
- Page 211 and 212: Figs. 93-96.-Texella cokendolpheri,
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- Page 215 and 216: Figs. 105-108.-Texelia mulaiki Good
- Page 217 and 218: and McCarty Caves, which are known
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- Page 221 and 222: Notes.-The type locality was errone
- Page 223 and 224: Figs. 128-131.-Taella reyesi, new s
- Page 225 and 226: Figs. 136-139.-Texella reyesi, new
- Page 227 and 228: Figs. 144-147.-Texella reyesi, new
- Page 229 and 230: Figs. 152-155.-Texella reyesi, new
- Page 231 and 232: 1989 (W. Elliott, J. Reddell, and M
- Page 233 and 234: Table 3.-Continued.# locality sex S
- Page 235 and 236: mesoapical; patella, 2 mesal; tibia
- Page 237 and 238: Figs. 162-165.-Texella gmbbsi, new
- Page 239 and 240: Figs. 166-169.-Texella diplospina,
- Page 241 and 242: Figs. 174-177.-Texella renkesae, ma
- Page 243 and 244: Figs. 178-18\.-Teulla spinoperca, n
- Page 245 and 246: Distribution.-Known only from Fayet
- Page 247 and 248: Figs. 190-193.-Texellafendi, new sp
- Page 249 and 250: CLASSIFICAnONTexellabifurcata group
- Page 251 and 252: Chandler, D.S. 1992. The Pselaphida
- Page 253 and 254: Key to Species1. Abdominal segments
- Page 255: stemite VI slightly impressed at ba
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- Page 261 and 262: small ventral carina near base, pro
- Page 263: Grigarick, A.A., and R.O. Schuster.
- Page 266 and 267: Cicurifla (Cicurella) holsiflgeri G