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searchable PDF - Association for Mexican Cave Studies

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inhabits tropical deciduous <strong>for</strong>est along the easternslopes of the Sierra de Guatemala in Tamaulipas; P.rowlandi occurs in the coastal plain east of CiudadValles in San Luis Potosi; P. occidentalis is the onlyspecies of Protoschizomidae known fromsouthwestern Mexico and the Pacific drainage.The cavernicole species of Protoschizomus appearto be less troglomorphic than the species ofAgastoschizomus, but are known only from caves.Protoschizomus gertschi inhabits a single cave nearMiquihuana in the western Sierra Madre Oriental. Itis separated from the species of the Purificaci6nregion by the Rio Guayalejo. The occurrence of fourspecies of Protoschizomidae in the Purificaci6nregion is remarkable. Agastoschizomus patd is ahighly troglomorphic species and is probablyrepresentative of a very early invasion of the cavesof this region. Agastoschizomus sp. cf. patei (knownonly from immature specimens) is sympatric inCueva del Tecolote with P. sprousei. The threespecies of Protoschizomus in the Purificaci6n regionare each known only from a single cave, but shouldbe found in other nearby caves with additionalcollecting. Protoschizomus sprousei occurs only inCueva del Tecolote near the village of Los SanPedro at an elevation of 1,450 m. The remainingtwo species, P. purijicacion and P. treacyae occurin Cueva X at an elevation of 1,950 m and in Cuevadel Borrego at an elevation of 1,980 m,respectively; both of these caves are located near thevillage of Conrado Castillo. The occurrence of threespecies of the same genus in the caves of this area isparalleled by the carabid beetle genusMexaphaenops Bolfvar y Pieltain with four speciesin the same area. Mexaphaenops jebriculosus Barr isfound at lower elevations only a few kilometersfrom Cueva del Tecolote. Three species, M.jamesoni Barr, M. mackenziei Barr and M.sulcifrons Barr inhabit caves in the Conrado Castilloarea. As with Mexaphaenops, the presence of threespecies of Protoschizomus in the same area mayrepresent different times of invasion of thecavernicole habitat. Protoschizomus purijicacionexhibits a lesser degree of troglobitic modification\j~19, 35,40, 43P. pnc/'ypnlplls group19,21,26,43vP. sprnlfse; group22,36Agnsrosc/'izofllfls17,24,27,34,39PrOlOschizomrlS25,2R,37Ilubbardiidae14-16,23,29,42Pro(oschizomidae1R, 20,31,32,38'--__---,,--Schizolllida0-13CaillarostomataFig. 2.-Cladogram of selected members of the superorder Camarostomata. See text <strong>for</strong> explanation and Tables 1 and 2 <strong>for</strong>character list (autapomorphies have been excluded) and presumed polarities.43

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