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SA with well developed prong and reduced carina(Figs. 105-108).Description.-Total body length, 1.49-2.21.Scute length, 1.31-1.49. Leg II length, 12.5-19.8.Leg II/Scute length, 12.5-15.3. (N = 20).Color of body yellowish orange, appendagesyellowish white. Body finely rugose; tuberclesabsent from eye mound, pars thoracica, tergitemargins, and coxae II-IV; coxa I with row of setatetubercles. Carapace without, or with 1 pair of verysmall, AT. Eye mound conical, reduced; retina andcornea absent. Palpal megaspines: trochanter, oneventral; femur, two mesoapical; patella, two mesal;tibia, two mesal. Tarsal count: 4-5 to 8-5 (or 6)-6(or 5).Male (Fern <strong>Cave</strong>): Total body length, 1.62.Scute length, 1.41; width, 1.31. Eye mound length,0.18; width, 0.23. Leg II length, 19.7. TrIV spurabsent. POP absent (Fig. 104).Penis (Figs. 102, 105-108): VPP with 2 dorsal,16 lateral, and 4 ventral setae; lateral setaemedium-sized, dorsal and ventral setae long; ASshort, curved, with trifurcate apex. Glans: BK along flap; 'ML broad with truncate apex; PSL arounded lobe lacking ventral notch and dorsal spine.S with apical half straight and thin; BF welldeveloped; SA with lateral prongs and reducedcarina, lacking serrations.Female (Fern <strong>Cave</strong>): Total body length, 1,69.Scute length, 1.44; width, 1.36. Eye mound length,0.21; width, 0.26. Leg II length, 18.5. TrIVlacking ventral tubercle.Ovipositor (Figs. 101, 109-112): cuticle intricatelyfolded; lacking microspines or tubercles;apical teeth absent.Variation.-The tarsal count is rather variable,especially in the tarsomere number of leg II wherealmost half of the specimens show variation fromthe typical number of -7-: one specimen has -5-,one -5/6- (-5- on the left leg, -6- on the right), one-617-, three -7/8-, and two -8-. In addition, twospecimens have counts of -5/6-, one on leg III theother on leg IV.The LII/SL varies from 12.5 to 15.3. The shortestleg lengths tend to occur in the southernpopulations (Boggus and Ezell's <strong>Cave</strong>s); the longestin the northern population (Flint Ridge <strong>Cave</strong>).Natural History.-T. mulaiki occurs sympatricallywith T. renkesae and T. diplospina butoccupies relatively deeper portions of the respectivecaves. These harvestmen appear to be relativelyuncommon, at least in Ezell's <strong>Cave</strong> where a 15­month faunal survey turned up only 7 specimens.On the other hand, we found a comparable numberof specimens during a few hours of collecting inFern <strong>Cave</strong>. That same study also suggests that theyare attracted to baits (6 specimens found at baitedrocks) and seem to prefer cheese (4 specimens)(Davis, 1979).Material Examined.-UNITED STATES:Texas: Hays Co.: [no specific locality], 15 Apr.1939 (S. Mulaik, AMNH), 1 female (type); SanMarcos, Ezell's <strong>Cave</strong>, 30 Apr. 1978 (J. Davis,WAS), 2 females; 2 July 1978 (1. Davis, WAS), 1female; Boggus <strong>Cave</strong>,. 16 Jan. 1988 (S. Harden,TMM), 2 females; McCarty <strong>Cave</strong>, 1988 [no specificdate] (A. Grubbs, TMM), 1 juvenile; McGlothlinSink, 26 May 1989 (A. Grubbs, J. Reddell, and M.Reyes, TMM), 1 juvenile; 3 Sep. 1989 (D. Ubick,S. Fend, and S. Renkes, CDU), 1 male, 3 juveniles;II mi W San Marcos, Fern <strong>Cave</strong>, 15 July 1988 (A.Grubbs, J. Evans, and L. Schneider, TMM), 1female; 26 May 1989 (A. Grubbs, J. Reddell, andM. Reyes, TMM), 1 female; 2 Sep. 1989 (D.Ubick, S. Fend, S. Renkes, and A. Grubbs, CDU,CAS), 3 males, 3 females; Ladder <strong>Cave</strong>, lowerlevel, 2 Sep. 1989 (D. Ubick, S. Fend, S. Renkes,J. Reddell, and M. Reyes, CDU), 1 female; 5 mi WKyle, Michaelis <strong>Cave</strong>, Jan. 1990 (A. Grubbs and L.Graves, TMM), 1 male. Travis Co.: Flint Ridge<strong>Cave</strong>, 8-9 June 1984 (D. Pate, TMM), 1 juvenile;Apr. 1989 (M. Grimm, TMM), 1 male; Apr. 1989(J. Reddell and M. Reyes, TMM), 1 juvenile; 25Nov. 1989 (M. Grimm, AMNH), 1 female, 1juvenile; Slaughter Creek <strong>Cave</strong>, 24 Feb.!3 Mar.1990 (W. Russell, TMM), 1 male; Whirlpool <strong>Cave</strong>,8 Jun. 1980 (S. Robertson, TMM), 1 juvenile.Distribution.-Known from the caves of CentralTexas, Hays and Travis Counties (Maps 2-5).Notes.-The sex of the holotype was originallystated as male but, as was subsequently corrected(Goodnight and Goodnight, 1967:7), is in fact afemale. All other specimens previously assigned tothis species actually represent anew, and not veryclosely related, species (T. reyesi).The type locality, although not specificallyindicated on the locality label, is probably Ezell's<strong>Cave</strong>, a popularly visited cave at the time S. Mulaikcollected the holotype (J. Reddell, personalcommunication). This is supported by the fact thatthe three recently collected female specimens fromEzell's <strong>Cave</strong> closely resemble the holotype.The present interpretation of T. mulaiki must beconsidered preliminary until male specimens fromEzell's <strong>Cave</strong> become available and are shown to beconspecific with the known males. Even moretentative is the inclusion of the two populations atthe extreme ends of the distribution, from Whirlpool203

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