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Horticulture Principles and Practices

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into the greenhouse or garden. Poor-quality sod often contains weed seedlings. Further,<br />

improperly laid sod with gaps between strips creates opportunities for weeds to rapidly<br />

infest a lawn.<br />

A plant’s life cycle or growth pattern influences the control strategy employed in<br />

weed control. Winter annuals are best controlled while still in the seedling stage during<br />

the fall <strong>and</strong> early spring. Similarly, summer annuals should be controlled soon after germination.<br />

Biennials are vegetative in the first year of growth; they should be controlled<br />

in that year. Perennials are very difficult to control <strong>and</strong> hence should not be allowed to<br />

become established. Once established, they have a persistent root system, making them<br />

difficult to control. Perennials can be controlled effectively during the period of rapid<br />

growth before flowering or during the regrowth period after cutting. Flowering species<br />

should be prevented from setting seed.<br />

Biological, chemical, <strong>and</strong> cultural control of weeds <strong>and</strong> other pests is discussed in<br />

Chapter 8. Annual weeds may be effectively controlled by mulching. However, mulching<br />

is ineffective in controlling perennial weeds. Controlling any type of weed is easier when<br />

the plants are in the seedling stages. Application of herbicides during the seedling stage<br />

is often effective in controlling weeds; however, it is ineffective when weeds are mature.<br />

Annuals <strong>and</strong> biennials also respond less favorably to chemical control when plants are<br />

entering the reproductive phase. In perennials, chemical control is effective at the bud<br />

stage, just before flowering.<br />

7.1.4 WEEDS AS INDICATORS OF SOIL FERTILITY<br />

Weeds may be plants out of place, but they arise where conditions are most favorable for<br />

their existence. The type of weed species found on a piece of l<strong>and</strong> is often a fairly good<br />

indicator of the soil characteristics (especially fertility, pH, <strong>and</strong> type). Examples of such<br />

telltale signs are as follows:<br />

1. An area of l<strong>and</strong> on which a good population of, for example, goosegrass, thistles,<br />

chickweed, <strong>and</strong> yarrow are found usually indicates that the soil is fertile <strong>and</strong><br />

nutritionally balanced.<br />

2. When d<strong>and</strong>elion, poppy, bramble, shepherd’s purse, bulbous buttercup, <strong>and</strong><br />

stinging nettle occur in dense populations, the soil is likely to be light <strong>and</strong> dry.<br />

3. Sedge, buttercup, primrose, thistle, dock, comfrey, <strong>and</strong> cuckooflower are found in<br />

wet soils.<br />

4. Acidic soils support acid-loving plants such as cinquefoil, cornflower, pansy,<br />

daisy, foxglove, <strong>and</strong> black bindweed.<br />

5. White mustard, bellflower, wild carrot, goat’s beard, pennycress, <strong>and</strong> horseshoe<br />

vetch are found in alkaline soils.<br />

6. Clay <strong>and</strong> heavy soils hold moisture <strong>and</strong> favor crops such as plantain, goosegrass,<br />

annual meadowgrass, <strong>and</strong> creeping buttercup.<br />

It should be emphasized that large populations of mixtures of several of the associated<br />

species listed must occur for the diagnosis to be reliable.<br />

7.2 PARASITIC PLANTS<br />

Parasitic Organism<br />

An organism that lives on<br />

or in another species <strong>and</strong><br />

derives part or all of its<br />

nourishment from the<br />

living host.<br />

There are more than 3,000 species of parasitic angiosperms. These plants have little or<br />

no chlorophyll <strong>and</strong> thus are incapable of photosynthesizing to meet their nutritional<br />

needs. Parasitic plants connect themselves to their host plants’ water- <strong>and</strong> foodconducting<br />

tissues through specialized rootlike projections called haustoria (singular:<br />

haustorium). The dodder (Cuscuta salina), a parasite with bright-yellow or orangecolored<br />

str<strong>and</strong>s, is a member of the family Convolvulaceae (morning glory family). It is<br />

a stem parasite. Others in this category are Loranthes <strong>and</strong> Arienthobium. Root parasites<br />

include the broomrapes (family Orobanchaceae). The Indian pipe (Monotropa uniflora)<br />

216 Chapter 7 Biological Enemies of Horticultural Plants

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