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PDF - Wallace Online

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78 ZOOLOGICAL GEO&RAPHY. [part iii.<br />

there is good reason to believe that many species have become<br />

extinct since the European occupation of them. When small<br />

islands are much cultivated, many of these molluscs which can<br />

only live under the shade of forests, are soon extirpated. In<br />

St. Croix many species have become extinct at a comparatively<br />

recent period, from the burning of forests ; and as we know that<br />

in all the islands many of the species are excessively local, being<br />

often confined to single valleys or ridges, we may be sure that<br />

wherever the native forests have disappeared before the hand of<br />

man, numbers of land-shells have disappeared with them. As<br />

some of the smaller islands have been almost denuded of their<br />

wood, and in the larger ones extensive tracts have been cleared<br />

for sugar cultivation, a very considerable number of species have<br />

almost certainly been exterminated.<br />

General Conclusions as to the Past History of the West Indian<br />

Islands.—The preceding sketch of the peculiarities of the animal<br />

life of these islands, enables us to state, that it represents the<br />

remains of an ancient fauna of decided Neotropical type, having<br />

on the whole most resemblance to that which now inhabits the<br />

Mexican sub-region. The number of peculiar genera in all<br />

classes of animals is so great in proportion to those in common<br />

with the adjacent mainland, as to lead us to conclude that,<br />

subsequent to the original separation from the Mexican area, a<br />

very large tract of land existed, calculated to support a rich and<br />

varied fauna, and, by the interaction of competing types, give<br />

rise to peculiar and specially modified organisms. We have<br />

already shown that the outline of the present islands and the<br />

depths of the surrounding seas, give indications of the position<br />

which not improbably occupied<br />

and extent of this ancient land ;<br />

the space enclosed by uniting Western Cuba with Yucatan, and<br />

Jamaica with the Mosquito Coast. This land must have<br />

stretched eastward to include Anguilla, and probably northward<br />

to include the whole of the Bahamas. At one time it perhaps<br />

extended southward so as to unite Hayti with northern<br />

Venezuela, while Panama and Costa Eica were sunk beneath the<br />

Pacific. At this time the Lesser Antilles had no existence.<br />

The only large island of whose geology we have any detailed

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