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PDF - Wallace Online

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CHAP. XV.] THE NEAKCTIC REGION. 117<br />

separation was effected by an arm of the sea across what is<br />

now Nicaragua, with perhaps anoth# at Panama. This would<br />

leave Mexico and Gimtemala joined to North America, and<br />

forming part of the Nearctic region, although no doubt contain-<br />

ing many Neotropical forms, which they had received during<br />

earlier continental periods ; and these countries might at other<br />

times have been made insular by a strait at the isthmus of<br />

Tehuantepec, and have then developed some peculiar species.<br />

The latest climatal changes have tended to restrict these<br />

Neotropical forms to those parts where the climate is really<br />

tropical; and thus Mexico has attained its present strongly<br />

marked Neotropical character, although deficient in many of<br />

the most important groups of that region.<br />

In view of these recent changes, it seems proper not to draw<br />

any decided line between the Nearctic and Neotropical regions,<br />

but rather to apply, in the case of each genus, a test which will<br />

show whether it was probably derived at a comparatively recent<br />

date fl'om one region or the other. The test referred to, is the<br />

existence of peculiar species of the genus, in what are un-<br />

doubtedly portions of ancient North or South America. If,<br />

for example, all the species of a genus occur in North America,<br />

some, or even all, o£ them, migrating into the Neotropical region<br />

in winter, while there are no peculiar NeolToj>ical species, then<br />

we must class that genus as strictly Nearctic; for if it were<br />

Neotropical it would certainly have developed some peculiar<br />

resident forms. Again, even if there should be one or two<br />

resident species peculiar to that part of Central America north<br />

of the ancient dividing strait, with an equal or greater number<br />

of species ranging over a large part of Temperate North America,<br />

the genus must still be considered Nearctic. Examples of the<br />

former case, are Helminthophaga and Myioclioctes, belonging to<br />

the Mniotiltidae, or wood-warblers, which range over all Tem-<br />

perate North America to Canada, where all the species are found,<br />

but in each case one of the species is found in South America,<br />

probably as a winter migrant. Of the latter, are Ammodramus<br />

soldi JiLuco (genera of finches), which range over the whole United<br />

States, but each have one peculiar species in Guatemala. These

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