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CHAP, xxiii.] SUMMARY AND CONCLUSION. .<br />

551<br />

organisms have spread over the earth, but owing to their small<br />

size and rapid multiplication, they have made use of some which<br />

are exclusively their own. Such are the passage along moun-<br />

tain ranges from the Arctic to the Antarctic regions, and the<br />

dispersal of certain types over all temperate lands. It will<br />

perhaps be found that insects have spread over the land surface<br />

in directions dependent on our surface zones—forests, pastures,<br />

and deserts ;—and a study of these, with a due consideration of<br />

the fact that narrow seas are scarcely a barrier to most of the<br />

groups, may assist us to understand many of the details of<br />

insect-distribution.<br />

Terrestrial Mollusca.<br />

The distribution of land-shells agrees, in some features, with<br />

that of insects, while in others the two are strongly contrasted.<br />

In both we see the effects of great antiquity, with some special<br />

means of dispersal ;<br />

but while in insects the general powers of<br />

motion, both voluntary and involuntary, are at a maximum, in<br />

land-molluscs they are almost at a minimum. Although to<br />

some extent dependent on vegetation and climate, the latter are<br />

more dependent on inorganic conditions, and also to a large<br />

extent on the general organic environment. The result of these<br />

various causes, acting through countless ages, has been to spread<br />

the main types of structure with considerable uniformity over<br />

the globe ; while generic and sub-generic forms are often<br />

wonderfully localized.<br />

Land-shells, even more than insects, seem, at first sight, to<br />

require regions of their own ; but we have already pointed out<br />

the disadvantages of such a method of study. It will be far<br />

more instructive to refer them to those regions and sub-regions<br />

which are found *to accord best with the distribution of the<br />

higher animals, and to consider the various anomalies they pre-<br />

sent as so many problems, to be solved by a careful study of<br />

their habits and economy, and especially by a search after the<br />

hidden causes which have enabled them to spread so widely<br />

over land and ocean.<br />

The lines of migration which land-shells have followed, can<br />

YoL. II.—36

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