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LOCALITY DATA. 66 specimens examined<br />

from AMNH, BMNH, CASC, CMNH, CNCI,<br />

DEIC, EMEC, FSCA, HAHC, MCZC, MGFT,<br />

MNHN, MXAL, OSAC, SMTD, USNM,<br />

ZMHB, ZMUH.<br />

COLO<strong>MB</strong>IA (60). ANTIOQUIA (5): Yarumal,<br />

No Data. CALDAS (1): Manizales. CAQUETÁ<br />

(1): No Data. CAUCA (25): País Pamba,<br />

Pereira, Popayán, Silvia (20 km W), Timbío,<br />

No Data. DISTRITO CAPITAL (3): Bogotá,<br />

Santé Fé de Bogotá. QUINDÍO (1): Quindío.<br />

VALLE DE CAUCA (10): Cali, Cali (18 km W),<br />

Calima, San Antonio. NO DATA (14).<br />

NO DATA (6).<br />

One specimen labeled “Barranquilla” was<br />

considered from Colombia with no further<br />

data. Four specimens labeled “Venezuela”<br />

were considered to have no data. One specimen<br />

labeled “Venezuela D. F. Co Narquata”<br />

was considered to have no data. These localities<br />

are either far out <strong>of</strong> the known range or<br />

too low in elevation to support populations <strong>of</strong><br />

this species.<br />

TEMPORAL DATA. January (1), February<br />

(1), March (3), April (3), May (2), July (8), September<br />

(2), October (2), December (1).<br />

21. Platycoelia humeralis Bates,<br />

1888<br />

(Figs. 59, 60)<br />

Platycoelia humeralis Bates, 1888 (valid<br />

name)<br />

CATALOG. Platycoelia humeralis, Bates<br />

1888:293 [original description]; Ohaus<br />

1904b:294, 297, 305, 306, 339, 340 [redescription];<br />

Ohaus 1905:167 [illustration]; Ohaus<br />

1918:177 [catalog listing]; Blackwelder<br />

1944:247 [checklist]; Machatschke 1965:56,<br />

58 [catalog listing]; De Ruette 1970:93 [catalog<br />

listing]; Machatschke 1972:302 [catalog<br />

listing]; Martínez 1976:328 [distribution];<br />

Thomas 1993:402 [checklist]; Morón<br />

1994b:17, 44 [distribution, biology, key to<br />

Scarabaeoidea <strong>of</strong> Sierra de Hidalgo, México];<br />

Morón 1995:195 [comment on distribution];<br />

Morón 1997:51 [redescription]; Ratcliffe<br />

MONOGRAPHIC REVISION OF PLATYCOELIA 97<br />

2002:26 [checklist]; Smith 2002a:381 [phylogenetic<br />

analysis <strong>of</strong> Brachysternina].<br />

TYPE SPECIMENS. Platycoelia humeralis<br />

Bates 1888: 293. Lectotype female at BMNH<br />

labeled a) “Type” (round label with red border,<br />

typeface), b) “Sp. figured.” (typeface), c) “V. de<br />

Chiriqui, 4000 ft. Champion.” (typeface), d)<br />

“Platycoelia humeralis Bates” (handwritten),<br />

e) “B. C. A. Coll. II. (2). Platycoelia humeralis.”<br />

(typeface), f) “PLATYCOELIA HUMERALIS<br />

BATES DET: A. B. T. SMITH 2001 LECTO-<br />

TYPE F” (red label, handwritten and typeface).<br />

Lectotype here designated. See<br />

Methods and Materials section for a statement<br />

<strong>of</strong> taxonomic purpose. Nine male paralectotypes<br />

at ANSP, BMNH, MCZC, MNHN (4),<br />

USNM, and ZMHB all bearing the label “V.<br />

de Chiriqui, 4000-6000 ft. Champion.” (typeface)<br />

and my yellow paralectotype label.<br />

Twenty-one female paralectotypes at AMNH,<br />

ANSP (2), BMNH (2), CNCI, DEIC (3), MCZC<br />

(2), MNHN (5), SMTD, USNM (2), and ZMHB<br />

(2) all bearing the same labels as above. One<br />

female paralectotype at MNHN labeled<br />

“Coatepec Mexico” (handwritten) and bearing<br />

my yellow paralectotype label. Five female<br />

paralectotypes at BMNH (2), USNM, and<br />

ZMHB (2) all bearing the label “Bugaba,<br />

Panama. Champion.” (typeface) and my yellow<br />

paralectotype label. Although Bugaba was<br />

not explicitly mentioned as a locality in the<br />

original description <strong>of</strong> the species, Bates<br />

(1888) must have used these specimens for his<br />

original description. Both Bugaba specimens<br />

from the BMNH bear the label “B. C. A. Coll.<br />

II. (2). Platycoelia humeralis.” This is a typical<br />

label seen on many specimens used to<br />

write the Biologia Centrali-Americana. Therefore<br />

these specimens are paralectotypes. The<br />

locations <strong>of</strong> approximately 13 remaining<br />

paralectotypes are unknown. Type locality:<br />

Volcán de Chiriqui, Chiriqui, Panamá.<br />

DESCRIPTION. Male (n=285). Length 21.5-<br />

29.3 mm, width 12.8-17.4 mm (Fig. 59). Color<br />

lime green (sometimes darkened to olive<br />

green or brownish-green) or yellow; elytral intervals<br />

1, 3, 5, 7 lighter green (sometimes obsolete,<br />

lighter yellow in yellow specimens);<br />

yellow elytron, pronotum margins; yellow

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