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MONOGRAPHIC REVISION OF PLATYCOELIA 147<br />

DIAGNOSIS. This species is distinguished<br />

from all other species in the genus Platycoelia<br />

by the following combination <strong>of</strong> characters:<br />

color cream or tan; frontoclypeal suture complete;<br />

apex <strong>of</strong> the labrum with a reduced, triangular<br />

tooth not overlapping the mentum;<br />

mentum with a medial notch; antenna 10-segmented;<br />

elytron without distinct longitudinal<br />

lines; elytral apices not capable <strong>of</strong> completely<br />

covering the pygidium; apex <strong>of</strong> the elytral suture<br />

without an acute spine; mesothoracic<br />

process nub to weakly projecting apically; apical<br />

abdominal spiracle strongly protuberant;<br />

mesotarsomere 5 and metatarsomere 5 without<br />

an internal tooth; parameres with the<br />

apex rounded, not expanded.<br />

DISTRIBUTION (Fig. 58). Andes Mountains<br />

from southern Colombia to southern Perú.<br />

Recorded from 1,800-4,000 m. Occurs in<br />

paramo and other grassland habitats.<br />

BIOLOGY. Smith and Paucar-Cabrera (2000)<br />

discussed the biology <strong>of</strong> P. lutescens. The<br />

adults emerge from the paramo when the winter<br />

rains begin (usually late October to early<br />

November). It has been observed that adult<br />

emergence coincides with rainy days and the<br />

sound <strong>of</strong> thunder (Whymper 1892, Onore<br />

1997). They emerge ‘en masse’ and have been<br />

seen by the thousands for a short period <strong>of</strong><br />

time. Whymper (1892) gives an excellent account<br />

<strong>of</strong> this in Travels Amongst the Great<br />

Andes <strong>of</strong> the Equator. On 14 February 1880<br />

while travelling to Volcán Cotopaxi (he was<br />

primarily on a mountain climbing expedition)<br />

Whymper writes “It was our intention to<br />

travel direct to Cotopaxi, but a violent storm<br />

drove us for refuge into Pedregal . . . at the<br />

northern foot <strong>of</strong> Rumiñahui.” The next day<br />

after the violent storm, Whymper continued<br />

toward Cotopaxi through the plain <strong>of</strong><br />

Limpiopungo, noting “I found here, in great<br />

numbers, a rather large beetle belonging to<br />

the same tribe as our cockchafer” (Whymper<br />

collected some specimens that were part <strong>of</strong><br />

the type series for Leucopelaea albescens [= P.<br />

lutescens]). Whymper also noted that the<br />

beetles were probably “emerging hindquarters<br />

first from the sandy soil.” The destruction<br />

<strong>of</strong> paramo habitat around Quito (for urban development)<br />

caused P. lutescens populations to<br />

decline in that area (Smith and Paucar-<br />

Cabrera 2000).<br />

LOCALITY DATA. 288 specimens examined<br />

from BCRC, BMNH, CASC, CMNH, DEIC,<br />

DJCC, FGIC, HAHC, HNHM, LACM, LEMQ,<br />

MCZC, MGFT, MLJC, MNHN, QCAZ, SMFD,<br />

SMTD, USNM, ZMHB, ZSMC.<br />

COLO<strong>MB</strong>IA (7). NARIÑO (7): Cumbal, Pasto.<br />

ECUADOR (272). BOLÍVAR (4): Cashca<br />

Totoras. CAÑAR (18): El Tambo. COTOPAXI<br />

(25): El Boliche, Latacunga, Limpiopungo,<br />

Parque Nacional Cotopaxi. ESMERALDAS<br />

(2): Mayronga. I<strong>MB</strong>ABURA (16): Juncal,<br />

Otavalo. PICHINCHA (162): Alóag, Cayambe<br />

(9.9 km NW), Chillogallo, Conocoto,<br />

Cumbayá, Ilaló, Machachi, Nono, Palmeras,<br />

Píntag, Quito, Santa Catalina, Tambillo, No<br />

data. TUNGURAHUA (4): Baños, Píllaro. NO<br />

DATA (37).<br />

PERÚ (5). CUSCO (1): Cusco. LIMA (1):<br />

Callanga. PUNO (1): No data. NO DATA (2).<br />

NO DATA (4).<br />

Doubtful locality labels include four specimens<br />

labeled “Guayquil, Ecuador,” two specimens<br />

labeled “Zarayaku Ecuador or.” and one<br />

specimen labeled Archidona. I considered<br />

these “Ecuador, no data” because these localities<br />

are too low in elevation to support populations<br />

<strong>of</strong> this species. One specimen labeled<br />

“Chili” is also doubtful (this locality is far out<br />

<strong>of</strong> the known range for the species) and I consider<br />

as a “No Data” specimen.<br />

TEMPORAL DATA. January (10), February<br />

(9), March (2), April (1), May (4), June (4),<br />

August (1), September (2), October (34) November<br />

(16), December (28).

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