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Platycoelia marginata and Platycoelia prasina]; Machatschke 1965:57 [catalog listing as synonym of Platycoelia marginata]; Machatschke 1972:301 [catalog listing as synonym of Platycoelia marginata]. Platycoelia scutellata, Guérin-Méneville 1855:585 [original description]; Harold 1869:1230 [catalog listing]; Ohaus 1904b:284, 338 [redescription]; Ohaus 1918:178 [catalog listing]; Blackwelder 1944:247 [checklist]; Ohaus 1952:9 [distribution]; Machatschke 1965:57 [catalog listing]; Machatschke 1972:301 [catalog listing]; Martínez 1976:328, 335 [redescription]; Martínez and Martínez 1994:545, 548 [comparison with Platycoelia selanderi]. Platycoelia levis (=Platycoelia laevis, lapsus calami), Harold 1869:1230 [catalog listing as synonym of Platycoelia prasina]; Ohaus 1904b:338 [checklist as synonym of Platycoelia marginata]. TYPE SPECIMENS. Platycoelia marginata Burmeister lectotype male at MLUH labeled a) “Merida” (handwritten), b) “PLATYCOE- LIA MARGINATA BURMEISTER M DET: A.B.T.SMITH 2001 LECTOTYPE” (red label, handwritten and typeface), c) “marginata * Quito Dup.” (green label, handwritten). Lectotype here designated. See Methods and Materials section for a statement of taxonomic purpose. One female paralectotype at MLUH labeled a) “PLATYCOELIA MARGINATA BURMEISTER F PARALECTOTYPE DET: A.B.T. SMITH 2001” (yellow label, handwritten and typeface). Burmeister (1844) did not state how many specimens were in the type series. The existence and location of other paralectotypes are unknown. Type locality: Mérida, Venezuela. Platycoelia olivacea Blanchard lectotype male at MNHN labeled a) “4 51” (round label, green on upperside, handwritten on underside), b) “P. olivacea. Cat. Mus. Brésil.” (green label, handwritten), c) “PLATYCOELIA OLIVACEA BLANCHARD LECTOTYPE A.B.T.SMITH 2001” (red label, handwritten and typeface), d) “Platycoelia marginata Burmeister, 1844 M Det:A.B.T.Smith 2002” (typeface). Lectotype here designated. See Methods and Materials section for a state- MONOGRAPHIC REVISION OF PLATYCOELIA 51 ment of taxonomic purpose. Blanchard (1851) did not explicitly state how many specimens were in the type series. The existence and location of paralectotypes are unknown. First placed in synonymy with P. marginata by Ohaus (1904b). Ohaus (1904b) discussed the variability (especially of color) of P. marginata when he placed P. olivacea in synonymy. After examination of the lectotype, I concur with his decision and maintain the syonymy. Type locality: Andes Mountains of northern South America (Blanchard [1851] stated “Brésil” in the original description but this is erroneous). Platycoelia laevis Burmeister lectotype male at MLUH labeled a) “PLATYCOELIA LAEVIS BURMEISTER LECTOTYPE A.B.T. SMITH 2001” (red label, handwritten and typeface), b) “laevis * Columb. Merida [illegible]” (green label, handwritten), c) “Platycoelia marginata Burmeister, 1844 M Det: A.B.T.Smith 2002” (typeface). Lectotype here designated. See Methods and Materials section for a statement of taxonomic purpose. One female paralectotype at MLUH labeled a) “PLATYCOELIA LAEVIS BUR- MEISTER F PARALECTOTYPE DET: A.B.T.SMITH 2001” (yellow label, handwritten and typeface), b) “Platycoelia marginata Burmeister, 1844 F Det:A.B.T. Smith 2002” (typeface). Burmeister (1855) did not state how many specimens were in the type series. The existence and location of other paralectotypes are unknown. First placed in synonymy with P. marginata by Ohaus (1904b). Ohaus (1904b) commented that Burmeister (1844) described P. laevis using a discolored, large specimen of P. marginata when he placed the two names in synonymy. After examination of the lectotype I concur with his decision and maintain the syonymy. Type locality: Mérida, Venezuela. Platycoelia scutellata Guérin-Méneville lectotype male at MNHN labeled a) “Scutellata Boliv Guérin Type” (handwritten), b) “Ex.Musæo Mniszech” (typeface), c) “Dr Ohaus Vidit 1903.” (typeface), d) “PLATY- COELIA SCUTELLATA GUÉRIN-MÉNE- VILLE LECTOTYPE A.B.T.SMITH 2001” (red label, handwritten and typeface), e) “Platycoelia marginata Burmeister, 1844 M
52 BULLETIN OF THE UNIVERSITYOF NEBRASKA STATE MUSEUM Det:A.B.T.Smith 2002” (typeface). Lectotype here designated. See Methods and Materials section for a statement of taxonomic purpose. Although the lectotype is labeled “Boliv,” this species does not occur in Bolivia. Gaetano Osculati collected the lectotype in the Napo region of Ecuador during his trip there in 1847 (Guérin-Méneville 1855; Papavero 1973). The description of P. scutellata by Guérin-Méneville (1855) did not explicitly state where the type series was collected. The purpose of that paper was to describe beetles collected by Gaetano Osculati (also spelled Cajetano Osculati) during his travels to Argentina, Chile, and Perú (1834-1836) and Ecuador and Brazil (1847-1848) (Guérin- Méneville 1855). Although the lectotype was labeled “Boliv,” this specimen was obviously collected on the eastern slopes of the Andes Mountains in Ecuador. This is evident from the distinct coloration of the abdominal sternites (dark brown medially, green laterally) and the large size (26.8 mm long) characteristic of P. marginata specimens collected in Napo, Ecuador. According to a published itinerary of Osculati’s trip (Papavero 1973); he collected around Baeza, Cosanga, and Archidona in Ecuador from 19 June – 28 July 1847. This is undoubtedly when the lectotype was collected. It is unknown how the lectotype got from Osculati to Mniszech (and ultimately to the MNHN); but at the time it was common for insect collectors to buy, sell, and trade specimens. The name P. scutellata was erroneously used for specimens of P. selanderi from southern Perú and Bolivia by several authors, including Ohaus (1904b), Martínez (1976), and Martínez and Martínez (1994). Platycoelia scutellata is a larger form of P. marginata that occurs on the eastern slope of the Andes Mountains. There are no consistent differences between the character states of the eastern Ecuadorian and other populations. Therefore these names are here synonymized. Guérin-Méneville (1855) did not explicitly state how many specimens were in the type series. The existence and location of paralectotypes are unknown. Type locality: in the vicinity of Baeza, Cosanga, and Archidona in Napo, Ecuador. NEW SYN- ONYMY. DESCRIPTION. Male (n=334). Length 17.8- 26.4 mm, width 9.5-14.6 mm. Color of dorsal surface olive green to lime green or yellow (sometimes darkened to brownish-green or dark yellow); pronotum, scutellum, elytron sometimes with yellow lateral margin; scutellum sometimes yellow. Ventral surface black to dark brown medially, sometimes olive green or yellowish-green laterally. Body ovate, convex. Head: Dorsal surface glabrous. Frons densely punctate, clypeus rugopunctate, punctures moderate. Frons not depressed. Frontoclypeal suture complete. Clypeal apex rounded. Eyes separated by approximately 3.0 transverse eye-widths. Labrum densely punctate, with moderately large, setose punctures, setae tawny. Apex of labrum with triangular, medial tooth, apex of tooth overlapping apex of mentum. Mandibular scissorial region with 1 tooth, molar region with strong lamellae. Maxilla with 3 cupshaped teeth. Mentum apically with medial tooth curved into oral cavity. Antenna 9-segmented; club approximately equal to other segments combined, approximately equal to length of frons. Pronotum: Surface glabrous, moderately to densely punctate, with small and moderate punctures. Marginal bead weak laterally, absent elsewhere. Elytron: Surface glabrous; longitudinal striae weakly impressed or not impressed, punctate; punctures moderate with dark coloration; intervals sparsely punctate, punctures small. Suture apically with acute spine (sometimes worn down). Pygidium: Width approximately 1.7 times length medially. Surface weakly convex, sparsely to moderately punctate; punctures moderately large to moderate, setose (near apex); setae short, tawny. Venter: Thorax moderately setose (glabrous medially), setae cream colored. Mesothoracic process projecting anteriorly past mesocoxa; shape conical, ventrally flattened. Abdomen sparsely setose to glabrous. Apical spiracles not extruding. Legs: Protibia with 1 apical tooth (second weak tooth sometimes visible). Mesotibia and metatibia widest medially. Protarsomeres 2-4 wider than long, cupshaped. Protarsomere 4 with internoapical stridulatory ridge. Protarsomere 5 with internomedial, stridulatory tooth. Meso-
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Bulletin of the University of Nebra
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C O N T E N T S Abstract...........
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LIST OF FIGURES Fig. 1. Platycoelia
Page 8 and 9: Bulletin of the University of Nebra
Page 10 and 11: Members of the genus Platycoelia De
Page 12 and 13: alone to divide up the rutelines (a
Page 14 and 15: eing the most recent authority on t
Page 16 and 17: els Amongst the Great Andes of the
Page 18 and 19: tion. Zischka was a German who live
Page 20 and 21: Puncture size: large = 0.17 mm or l
Page 22 and 23: 26. Pronotal surface sparsely punct
Page 24 and 25: Fig. 11. Platycoelia alternans male
Page 26 and 27: MONOGRAPHIC REVISION OF PLATYCOELIA
Page 28 and 29: Fig. 33. Platycoelia rufosignata me
Page 30 and 31: CNCI Canadian National Collection o
Page 32 and 33: subspecies AND varieties in the sam
Page 34 and 35: PHYLOGENETIC PATTERNS AND BIOGEOGRA
Page 36 and 37: metaunguitractor plate usually with
Page 40 and 41: larvae, and Paucar-Cabrera and Smit
Page 50 and 51: Erichson (1847) did not state how m
Page 52 and 53: species]; Ohaus 1918:178 [catalog l
Page 60 and 61: tarsomere and metatarsomere 5 with
Page 62 and 63: 5. Platycoelia meridensis new speci
Page 64 and 65: in apical half. Protarsomere 4 with
Page 68 and 69: 7. Platycoelia prasina Erichson, 18
Page 70 and 71: written and typeface), e) “PLATYC
Page 72 and 73: mentum. Apex of mentum with triangu
Page 74 and 75: 1972:302 [catalog listing as subspe
Page 76 and 77: green to yellow; parameres with the
Page 82 and 83: duced tooth. Apex of mentum with tr
Page 84 and 85: Mnisnech,” “Dr Ohaus Vidit 1903
Page 86 and 87: VARIATION. Male (n=19). Length 19.2
Page 88 and 89: “Platycoelia forcipalis cotype Oh
Page 90 and 91: males with internomedial stridulato
Page 94 and 95: tures moderate with dark coloration
Page 96 and 97: gins. Ventrally with dark brown met
Page 98 and 99: 19. Platycoelia flavostriata (Latre
Page 102 and 103: 20. Platycoelia nigrosternalis Ohau
Page 104 and 105: LOCALITY DATA. 66 specimens examine
Page 108 and 109:
MONOGRAPHIC REVISION OF PLATYCOELIA
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Calloodes nitidissimus Lea, 1919 (C
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