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MONOGRAPHIC REVISION OF PLATYCOELIA 105 an internobasal tooth; parameres with the apex dorsoventrally flattened, without a setose patch. DISTRIBUTION (Fig. 63). Known only from Tatamá, Colombia. LOCALITY DATA. 1 specimen examined from BMNH. COLOMBIA (1). RISARALDA (1): Tatamá. 25. Platycoelia variolosa Ohaus, 1904 (Fig. 63) Platycoelia variolosa Ohaus, 1904 (valid name) CATALOG. Platycoelia variolosa, Ohaus 1904b:304, 305, 339 [original description]; Ohaus 1918:178 [catalog listing]; Blackwelder 1944:247 [checklist]; Machatschke 1965:58 [catalog listing]; Machatschke 1972:302 [catalog listing]. TYPE SPECIMENS. Platycoelia variolosa Ohaus lectotype male at ZMHB labeled a) “COLOMBIE” (light green label, typeface), b) “Platycoelia variolosa Type Ohs.” (orange label, handwritten), c) “PLATYCOELIA VARIOLOSA OHAUS M LECTOTYPE A.B.T.SMITH 2002” (red label, handwritten and typeface). Lectotype here designated. See Methods and Materials section for a statement of taxonomic purpose. One female paralectotype at ZMHB labeled a) “COLOMBIE” (light green label, typeface), b) “F” (typeface), c) “Platycoelia variolosa Cotype Ohs.” (orange label, handwritten), d) “PLATY- COELIA VARIOLOSA OHAUS F PARA- LECTOTYPE A.B.T.SMITH 2002” (yellow label, handwritten and typeface). Two male paralectotypes at MNHN labeled a) “Colum” (handwritten), b) “Ex-Musæo D.Sharp 1890” (typeface), c) “Dr Ohaus Vidit 1903.” (typeface), d) “Ohaus determ. variolosa Ohaus M cotyp.” (typeface and handwritten), e) “PLATYCOELIA VARIOLOSA OHAUS M PARALECTOTYPE A. B. T. SMITH 2002” (handwritten and typeface). One male paralectotype at MNHN labeled a) “Ex- Musæo E.Steinheil” (typeface), b) “Dr Ohaus Vidit 1903.” (typeface), c) “Ohaus determ. Pl. variolosa Ohaus M cotyp.” (typeface and handwritten), e) “PLATYCOELIA VARIOLOSA OHAUS M PARALECTOTYPE A.B.T. SMITH 2002” (handwritten and typeface). One Ecuadorian specimen of P. paucarae from ZMHB bears Ohaus’ orange type label stating “Platycoelia variolosa cotype Ohs.” Another Ecuadorian specimen (of P. nigrosternalis) at NMPC bears Ohaus’ determination label stating “Platycoelia variolosa Cotype F Ohs.” Since Ecuador was not mentioned in the original description of P. variolosa, neither of these specimens are part of the type series. Type locality: Colombia. DESCRIPTION. Male (n=10). Length 26.8- 29.1 mm, width 14.4-17.3 mm. Color dorsally light green to olive green with yellow elytron, pronotum, and scutellum margins. Ventrally light green with metasternum, abdominal sternites black medially (sometimes black extends to lateral margins). Body ovate, strongly convex. Head: Dorsal surface glabrous. Frons densely punctate, clypeus rugopunctate, punctures small or moderate. Frons not depressed. Frontoclypeal suture incomplete, absent medially. Clypeus subrectangular. Eyes separated by approximately 3.0 transverse eye-widths. Labrum moderately punctate, with moderately large, setose punctures, setae tawny. Apex of labrum with triangular, medial tooth; apex of tooth broadly truncate, extending past apex of mentum. Apex of mentum with triangular medial tooth curved into oral cavity. Antenna 10-segmented; club approximately equal to other segments combined, slightly longer than length of frons. Pronotum: Surface glabrous, densely punctate, with small or moderate punctures. Marginal bead weak apicolaterally, absent elsewhere. Elytron: Surface glabrous; longitudinal striae weakly punctate, punctures moderate with dark coloration; intervals densely punctate, punctures small. Suture angled apically, without spine. Pygidium: Width approximately 2.7 times length medially. Surface weakly convex, densely punctate; punctures small or moder-
106 BULLETIN OF THE UNIVERSITYOF NEBRASKA STATE MUSEUM ate, setose; setae short, cream colored. Venter: Thorax densely setose (except glabrous medially), setae white. Mesothoracic process strongly produced, projecting anteriorly past protrochanter; shape conical-elongate, apex rounded, dorsoventrally flattened. Abdomen sparsely setose, setae white. Apical spiracles not extruding. Legs: Protibia with 3 teeth in apical half; apical 2 teeth longer, third tooth short, obsolete. Mesotibia and metatibia widest medially. Protarsomeres 2-4 wider than long, cup-shaped. Protarsomere 3-4 with internoapical stridulatory ridge. Protarsomere 5 with internomedial stridulatory tooth. Mesotarsomere and metatarsomere 5 without internal tooth. Prounguitractor plate cylindrical, with 2 subapical setae. Mesounguitractor plate and metaunguitractor plate cylindrical, with 2 apical setae. Modified protarsal claw with length approximately equal to protarsomere 5, thickened and elongate when compared with other claw, laterally flattened, apex unevenly bifurcate with 1 bifurcation twisted beneath the other in ventral view. Modified mesotarsal and metatarsal claws elongated with ventral tooth, not thickened. Male Genitalia: Phallobase 1.4 times longer than length of parameres. Parameres dorsoventrally flattened, apicomedially acute. Female (n=10). Length 27.2-30.9 mm, width 14.8-16.8 mm. As male except in the following respects. Head: Antennal club approximately equal to segments 2-7. Legs: Protarsomere 3-4 without internoapical stridulatory ridge. Protarsomere 5 without internal tooth. Modified protarsal claw with ventral tooth, not thickened. DIAGNOSIS. This species is distinguished from all other species in the genus Platycoelia by the following combination of characters: frontoclypeal suture incomplete; labrum with a well-developed, truncate apical tooth strongly overlapping the mentum; mentum with a distinct tooth curved into the oral cavity; antenna 10-segmented; pronotum surface densely punctate with a mix of small and moderate punctures; elytron without distinct, yellow, longitudinal lines; elytral apices capable of completely covering the pygidium; elytron with a distinct yellow suture; pygidium entirely setose; metasternum and abdominal sternites green or yellow; protibial apical 2 teeth adjacent, located in the apical quarter of the protibia; protibial spur noticeably shorter than the adjacent tibial apex; protarsomere 3 in males without a distinct apical stridulatory patch; modified protarsal claw in males laterally flattened; mesotarsomere 4 and metatarsomere 4 in males with an apical stridulatory ridge; mesotarsomere 5 and metatarsomere 5 with an internobasal tooth; parameres with the apex dorsoventrally flattened. DISTRIBUTION (Fig. 63). Andes Mountains in central Colombia. LOCALITY DATA. 20 specimens examined from AMNH, CASC, DEIC, MNHN, VMCP, ZMHB, ZMUH. COLOMBIA (19). ANTIOQUIA (2) Medellín. BOYACÁ (1): Villa de Leiva. CUNDINAMARCA (4): Fusagasugá, Monterredondo, Viotá. DISTRITO CAPI- TAL (1): Bogotá. NO DATA (11). NO DATA (1). TEMPORAL DATA. June (1). 26. Platycoelia grandicula new species (Figs. 2, 8, 13, 61, 62) TYPE SPECIMENS. Male holotype, female allotype and 92 paratypes (58 male, 34 female). Holotype male at UNSM labeled “PANAMA, PANAMA PR. 10 KM N EL LL- ANO MAY 16-22 1987 E. GIESBERT, COLL.” and “Donated by Ed Giesbert 1997.” Allotype female at UNSM labeled “PANAMA, PANAMA PR. ±10 KM N EL LLANO, 1400’ MAY 16-20 1984 E. GIESBERT, COLL.” and “Donated by Ed Giesbert 1997.” One male paratype at FSCA labeled “PANAMA, PANAMA PR. 10-12 KM N EL LLANO JUNE 3-8 1986 E. GIESBERT, COLL.” Two male paratypes at DCCC labeled “PANAMA Panama Pr. El Llano-Cartí Rd. 8 Km N El Ll-
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Bulletin of the University of Nebra
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C O N T E N T S Abstract...........
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LIST OF FIGURES Fig. 1. Platycoelia
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Bulletin of the University of Nebra
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Members of the genus Platycoelia De
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alone to divide up the rutelines (a
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eing the most recent authority on t
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els Amongst the Great Andes of the
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tion. Zischka was a German who live
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Puncture size: large = 0.17 mm or l
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26. Pronotal surface sparsely punct
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Fig. 11. Platycoelia alternans male
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MONOGRAPHIC REVISION OF PLATYCOELIA
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Fig. 33. Platycoelia rufosignata me
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CNCI Canadian National Collection o
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subspecies AND varieties in the sam
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PHYLOGENETIC PATTERNS AND BIOGEOGRA
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metaunguitractor plate usually with
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larvae, and Paucar-Cabrera and Smit
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Erichson (1847) did not state how m
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species]; Ohaus 1918:178 [catalog l
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Platycoelia marginata and Platycoel
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tarsomere and metatarsomere 5 with
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Page 66 and 67: MONOGRAPHIC REVISION OF PLATYCOELIA
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Page 86 and 87: VARIATION. Male (n=19). Length 19.2
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Calloodes nitidissimus Lea, 1919 (C
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