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MONOGRAPHIC REVISION OF PLATYCOELIA 177 obscured by punctures. Clypeus broadly rounded, apex reflexed. Eyes separated by approximately 4.3 transverse eye-widths. Labrum rugose, setose; setae tawny. Apex of labrum with triangular, medial tooth, apex of tooth well separated from apex of mentum. Mandibular scissorial region with 1 tooth, molar region smooth. Maxilla with 3 apical teeth. Apex of mentum with medial notch. Antenna 9-segmented; club slightly shorter than other segments combined, approximately equal to length of frons. Pronotum: Surface glabrous, sparsely punctate, with moderate punctures. Marginal bead present apically (except medially) and laterally, absent basally. Elytron: Surface glabrous; longitudinal striae weakly impressed, impunctate to weakly punctate; intervals impunctate. Suture apically rounded. Pygidium: Width approximately 1.9 times length medially. Surface smooth, weakly convex, impunctate (except sparsely punctate near apex); punctures moderate, setose; setae long, tawny. Venter: Thorax densely setose; setae long, tawny. Mesothoracic process barely produced, projecting anteriorly to subapex of mesocoxa. Abdomen sparsely setose, setae tawny. Apical spiracles not extruding. Legs: Protibia with 3 teeth in apical half; first and second teeth subequal in size; third tooth short, removed slightly from apical teeth. Mesotibia and metatibia widest apically. Protarsomeres 2-4 wider than long, cup-shaped. Protarsomere 4 with weak internoapical stridulatory ridge. Protarsomere 5 with internomedial stridulatory tooth. Mesotarsomere 5 and metatarsomere 5 without internal tooth. Unguitractor plate cylindrical, with 1 apical and 1 subapical seta. Modified protarsal claw with length slightly shorter than protarsomere 5, thickened and elongated when compared with other claw, flattened on slightly diagonally plane, apex unevenly bifurcate. Modified mesotarsal and metatarsal claws elongated with ventral tooth, not thickened. Male Genitalia: Phallobase 1.5 times longer than length of parameres. Parameres with apex rounded. Female: Unknown. DIAGNOSIS. This species is distinguished from all other species in the genus Platycoe- lia by the following combination of characters: dorsal color black with light yellowish-brown elytron; frons sparsely setose laterally; frontoclypeal suture complete; apex of the labrum with a reduced, triangular tooth not overlapping the mentum; mentum with a medial notch; antenna 10-segmented; elytron without distinct, yellow, longitudinal lines; elytral apices not capable of completely covering the pygidium; apex of the elytral suture without an acute spine; mesothoracic process nub to weakly projecting apically; apical abdominal spiracle not protuberant; mesotarsomere 5 and metatarsomere 5 without an internal tooth; parameres with the apex rounded, not expanded. DISTRIBUTION (Fig. 73). Andes Mountains in La Paz, Bolivia. LOCALITY DATA. 4 specimens examined from USNM, ZMHB. BOLIVIA (4). LA PAZ (4): Chulumani, Pongo de Quime. TEMPORAL DATA. July (1), October (3). 59. Platycoelia alticola (Gutiérrez, 1951) (Fig. 73) Callichloris (Epicallichloris) alticola Gutiérrez, 1951 (original combination) CATALOG. Callichloris (Epicallichloris) alticola, Gutiérrez 1951:112, 115, 116 [cited as type species for Callichloris (Epicallichloris), original description, key to species of Callichloris]; Gutiérrez 1952:226 [comparison with Callichloris haenkei]. Platycoelia alticola, Machatschke 1965:60 [new combination, catalog listing]; Machatschke 1972:304 [catalog listing]; Smith and Jameson 2001:105 [comparison with Eremophygus bicolor]. TYPE SPECIMENS. Callichloris (Epicallichloris) alticola Gutiérrez neotype male at FMNH labeled a) “Bolivia 10-IX-49 Cochabamba 4000 ms – Pena coll” (handwrit-
178 BULLETIN OF THE UNIVERSITYOF NEBRASKA STATE MUSEUM ten), b) “Callichloris (Epicallichloris) alticola – Gut M R. Gutiérrez-Det.50” (handwritten and typeface), c) “CALLICHLORIS ALTI- COLA GUTIÉRREZ M PARATYPE” (yellow label, handwritten and typeface), d) “FMNH, 1986 L. Peña Coll. Acc. # 17-422” (typeface), e) “CALLICHLORIS ALTICOLA GUTIÉR- REZ NEOTYPE A.B.T.SMITH 2002” (red label, handwritten and typeface). f) “Platycoelia alticola (Gutiérrez, 1951) M Det:A.B.T.Smith 2002” (typeface). The location of the holotype from Gutiérrez’s original type series is unknown. The holotype male was deposited in Antonio Martínez’s personal collection (Gutiérrez 1951). This collection was purchased by Henry and Anne Howden (HAHC), and at the time of purchase, all the primary types were deposited at MACN. The holotype is not at MACN according to Axel Bachmann (a curator at MACN) and Federico Ocampo (who searched for the specimen on my behalf), HAHC (which I searched myself), or any of the other collections from which I borrowed material. I can only conclude that it has been lost. The neotype is also the sole paratype of this species (Gutiérrez 1951). Gutiérrez based the original description on two specimens. The holotype was from Bolivia with the following data “Chapare dep. de Cochabamba; Yungas del Palmar; 3500 m; I-1949” (Gutiérrez 1951). The neotype is also from Cochabamba (Bolivia), therefore, the type locality for the species is not significantly altered. Neotype here designated. See Methods and Materials section for a statement of taxonomic purpose. The neotype was originally deposited as a paratype by Gutiérrez in the “Sociedad Científica Chilena Claudio Gay” collection. This short-lived collection was absorbed into the Luis Peña Collection, which was purchased by FMNH in 1986. Type locality: Cochabamba, Bolivia. DESCRIPTION. Male (n=2). Length 15.1- 16.6 mm, width 8.9-9.1 mm. Color dorsally dark brown to black with light brown elytron; ventrally brown to light brown. Body ovate, convex. Head: Dorsal surface setose, setae tawny. Frons moderately punctate (base) to rugose (apex), clypeus rugose, punctures moderate. Frons slightly depressed. Frontoclypeal suture obscured by punctures. Clypeus broadly rounded, apex reflexed. Eyes separated by approximately 4.2 transverse eyewidths. Labrum rugose, setose; setae tawny. Apex of labrum with triangular, medial tooth, apex of tooth well separated from apex of mentum. Apex of mentum with weak medial notch. Antenna 9-segmented; club slightly longer than other segments combined, slightly longer than frons. Pronotum: Surface setose along apex; setae long, tawny. Surface moderately punctate, with moderate punctures. Marginal bead present apically (except medially), laterally, absent basally. Elytron: Surface glabrous; longitudinal striae weakly punctate; impunctate. Suture apically angled. Pygidium: Width approximately 2.0 times length medially. Surface rugose, weakly convex, moderately setose; setae long, tawny. Venter: Thorax densely setose; setae long, tawny. Mesothoracic process reduced to nub, adjacent to mesocoxa. Abdomen sparsely setose, setae tawny. Apical spiracles not extruding. Legs: Protibia with 3 subequal in size teeth in apical half. Mesotibia and metatibia widest medially, apically. Protarsomeres 2-4 wider than long, cup-shaped. Protarsomere 4 with weak internoapical stridulatory ridge. Protarsomere 5 without internal tooth. Mesotarsomere 5 and metatarsomere 5 without internal teeth. Unguitractor plate cylindrical, with 1 apical and 1 subapical seta. Modified protarsal claw with length approximately equal to protarsomere 5, thickened and elongated when compared with other claw, flattened dorsoventrally, apex unevenly bifurcate (small bifurcation reduced to nub). Modified mesotarsal claw elongated, apex unevenly bifurcate. Modified metatarsal claw elongate, with ventral tooth, not thickened. Male Genitalia: Phallobase 1.2 times longer than length of parameres. Parameres with apex rounded, weakly expanded subapically. Female: Unknown. DIAGNOSIS. This species is distinguished from all other species in the genus Platycoelia by the following combination of characters: dorsal color dark brown to black with light brown elytron; frons sparsely setose laterally; frontoclypeal suture complete; apex of the la-
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Bulletin of the University of Nebra
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C O N T E N T S Abstract...........
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LIST OF FIGURES Fig. 1. Platycoelia
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Bulletin of the University of Nebra
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Members of the genus Platycoelia De
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alone to divide up the rutelines (a
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eing the most recent authority on t
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els Amongst the Great Andes of the
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tion. Zischka was a German who live
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Puncture size: large = 0.17 mm or l
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26. Pronotal surface sparsely punct
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Fig. 11. Platycoelia alternans male
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MONOGRAPHIC REVISION OF PLATYCOELIA
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Fig. 33. Platycoelia rufosignata me
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CNCI Canadian National Collection o
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subspecies AND varieties in the sam
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PHYLOGENETIC PATTERNS AND BIOGEOGRA
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metaunguitractor plate usually with
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larvae, and Paucar-Cabrera and Smit
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Erichson (1847) did not state how m
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species]; Ohaus 1918:178 [catalog l
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Platycoelia marginata and Platycoel
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tarsomere and metatarsomere 5 with
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5. Platycoelia meridensis new speci
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in apical half. Protarsomere 4 with
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7. Platycoelia prasina Erichson, 18
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written and typeface), e) “PLATYC
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mentum. Apex of mentum with triangu
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1972:302 [catalog listing as subspe
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green to yellow; parameres with the
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duced tooth. Apex of mentum with tr
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Mnisnech,” “Dr Ohaus Vidit 1903
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VARIATION. Male (n=19). Length 19.2
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“Platycoelia forcipalis cotype Oh
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males with internomedial stridulato
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tures moderate with dark coloration
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gins. Ventrally with dark brown met
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19. Platycoelia flavostriata (Latre
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20. Platycoelia nigrosternalis Ohau
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LOCALITY DATA. 66 specimens examine
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Page 134 and 135: MONOGRAPHIC REVISION OF PLATYCOELIA
Page 206 and 207: Calloodes nitidissimus Lea, 1919 (C
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