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32 BULLETIN OF THE UNIVERSITYOF NEBRASKA STATE MUSEUM<br />

Arrow 1899:369 [comment on morphology];<br />

Ohaus 1904b:258, 273, 277, 307, 332, 339 [redescription,<br />

species list]; Ohaus 1905:120<br />

[comparison with Leucopelaea]; Ohaus<br />

1918:178 [catalog listing]; Blackwelder<br />

1944:247 [checklist]; Gutiérrez 1951:112, 113,<br />

115 [redefinition <strong>of</strong> genus]; Ohaus 1952:9 [catalog<br />

listing]; Machatschke 1965:13, 55 [distribution,<br />

placed in synonymy with Platycoelia];<br />

Machatschke 1972:301 [catalog listing as synonym<br />

<strong>of</strong> Platycoelia]; Martínez 1976:327 [distribution];<br />

Morón 1997:50 [comment on<br />

nomenclature]; Smith and Jameson 2001:104<br />

[comment on nomenclature]; Ratcliffe and<br />

Ocampo 2002:368 [comment on nomenclature].<br />

Leucopelaea, Bates 1891a:4 [distribution];<br />

Bates 1891b:30 [original description];<br />

Whymper 1892:137 [biology]; Arrow 1899:369<br />

[comparison with Callichloris]; Ohaus 1904b:<br />

258, 277 [comment on morphology]; Ohaus<br />

1905:120, 164 [redescription, species list];<br />

Ohaus 1918:178 [catalog listing]; Blackwelder<br />

1944:247 [checklist]; Machatschke 1965:13, 55<br />

[distribution, placed in synonymy with<br />

Platycoelia]; Machatschke 1972:301 [catalog<br />

listing as synonym <strong>of</strong> Platycoelia]; Martínez<br />

1976:327 [distribution]; Morón 1997:50 [comment<br />

on nomenclature]; Smith and Paucar-<br />

Cabrera 2000:412 [comment on classification];<br />

Smith and Jameson 2001:104 [comment on nomenclature].<br />

Callichloris (Callichloris), Gutiérrez<br />

1951:115 [redescription as subgenus, key to<br />

species].<br />

Callichloris (Epicallichloris), Gutiérrez<br />

1951:115 [original description <strong>of</strong> subgenus,<br />

key to species]; Smith and Jameson 2001:104<br />

[comment on nomenclature].<br />

Epicallichloris, Machatschke 1965:60<br />

[placed in synonymy with Platycoelia];<br />

Machatschke 1972:304 [catalog listing as synonym<br />

<strong>of</strong> Platycoelia].<br />

TYPE SPECIES. Platycoelia Dejean, 1833.<br />

Type species Melolontha flavostriata Latreille,<br />

1813 by monotypy. Gender: feminine.<br />

Callichloris Burmeister, 1844. Type species<br />

Callichloris signaticollis Burmeister,<br />

1844 by monotypy. Gender: feminine.<br />

Leucopelaea Bates, 1891. Type species<br />

Leucopelaea albescens Bates, 1891 by<br />

monotypy. Gender: feminine.<br />

Epicallichloris Gutiérrez, 1951. Type species<br />

Callichloris alticola Gutiérrez, 1951 by<br />

original designation. Epicallichloris was<br />

originally described as a subgenus <strong>of</strong> Callichloris.<br />

Gender: feminine.<br />

DESCRIPTION. Length 10.5-34.0 mm,<br />

width 6.2-19.3 mm. Color usually lime green<br />

to olive green or yellow, sometimes dark green<br />

or brown to black. Body usually ovate,<br />

strongly to weakly convex. Head: Dorsal surface<br />

punctate, usually glabrous. Clypeal apex<br />

rounded to trapezoidal. Frontoclypeal suture<br />

complete or obsolete medially. Labrum projecting<br />

ventrally, perpendicular to clypeus.<br />

Apex <strong>of</strong> labrum with medial tooth. Mentum<br />

apically with notch or medial tooth. Antenna<br />

9-segmented or 10-segmented. Pronotum:<br />

Surface usually glabrous, always punctate.<br />

Marginal bead weak laterally, usually absent<br />

elsewhere. Elytron: Surface glabrous; longitudinal<br />

striae impressed or not impressed,<br />

punctate or impunctate. Suture apically<br />

rounded, angled, or with acute spine. Pygidium:<br />

Surface convex, punctate, setose. Venter:<br />

Thorax setose. Mesothoracic process<br />

present, <strong>of</strong>ten projecting anteriorly to<br />

procoxa. Abdomen sparsely setose or glabrous.<br />

Apical spiracles sometimes extruding.<br />

Legs: Protibia with 1-3 teeth. Protarsomeres 2-<br />

4 wider than long, cup-shaped. Protarsomeres<br />

3-4 in males <strong>of</strong>ten with internoapical stridulatory<br />

ridge. Protarsomere 5 in males <strong>of</strong>ten with<br />

internomedial, stridulatory tooth. Unguitractor<br />

plate cylindrical with 2 setae. Modified<br />

protarsal claw in males thickened, elongate<br />

when compared with other claw, apex unevenly<br />

bifurcate. Modified protarsal claw in<br />

females with ventral tooth, not thickened.<br />

Modified mesotarsal claw weakly thickened,<br />

elongate when compared with other claw,<br />

apex unevenly bifurcate or with ventral tooth,<br />

not thickened. Modified metatarsal claw elongate<br />

with ventral tooth, not thickened. Male<br />

Genitalia: Phallobase not fused with<br />

parameres. Parameres not fused but longitudinally<br />

contiguous. Female Genitalia: Generally<br />

not diagnostic.<br />

LARVAE. Our knowledge <strong>of</strong> Platycoelia larvae<br />

is scant. Ohaus (1908, 1909a) published<br />

some rudimentary observations <strong>of</strong> P. gaujoni

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