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C O N T E N T S Abstract..............................................................................................................1 Introduction........................................................................................................3 Tribe Anoplognathini........................................................................................4 Subtribe Platycoeliina......................................................................................6 Taxonomic History of the genus Platycoelia......................................................6 Notes on Authors, Types, and Collections..........................................................8 Phylogenetic Methods and Materials...............................................................12 Phylogenetic Analysis................................................................................12 Character Definition..................................................................................12 Adult Morphological Character Analysis..................................................13 Phylogenetic Protocol.................................................................................22 Taxonomic Methods and Materials..................................................................22 Specimens.................................................................................................22 Catalog Format............................................................................................24 Type Specimens...........................................................................................24 Designation of Lectotypes and Neotypes....................................................25 Species Concept..........................................................................................26 Phylogenetic Patterns and Biogeographic Implications...................................27 Key to the Neotropical Genera of Anoplognathini...........................................30 Subtribe Platycoeliina Burmeister, 1844...........................................................31 Genus Platycoelia Dejean, 1833.........................................................................31 Key to the Species of Platycoelia........................................................................34 1. Platycoelia alternans Erichson, 1847..................................................42 2. Platycoelia inflata Ohaus, 1904..........................................................44 3. Platycoelia nervosa Kirsch, 1871........................................................48 4. Platycoelia marginata Burmeister, 1844............................................50 5. Platycoelia meridensis new species.....................................................55 6. Platycoelia selanderi Martínez and Martínez, 1994...........................57 7. Platycoelia prasina Erichson, 1847......................................................61 8. Platycoelia simplicior Ohaus, 1909.....................................................62 9. Platycoelia abdominalis Ohaus, 1904.................................................64 10. Platycoelia pomacea Erichson, 1847...................................................66 11. Platycoelia occidentalis Ohaus, 1904..................................................69 12. Platycoelia valida Burmeister, 1844....................................................71 13. Platycoelia mesosternalis Ohaus, 1904...............................................74 14. Platycoelia peruviana new species......................................................76 15. Platycoelia forcipalis Ohaus, 1904......................................................80 16. Platycoelia sandia new species...........................................................84 17. Platycoelia penai Frey, 1967...............................................................86 18. Platycoelia galerana new species........................................................87 19. Platycoelia flavostriata (Latreille, 1813).............................................91 20. Platycoelia nigrosternalis Ohaus, 1904..............................................95 21. Platycoelia humeralis Bates, 1888.....................................................97 22. Platycoelia intermedia Ohaus, 1925....................................................101 iii
23. Platycoelia convexa new species.......................................................102 24. Platycoelia interstincta new species.................................................104 25. Platycoelia variolosa Ohaus, 1904......................................................105 26. Platycoelia grandicula new species...................................................106 27. Platycoelia hiporum new species.......................................................111 28. Platycoelia paucarae new species......................................................112 29. Platycoelia flavoscutellata Ohaus, 1904............................................115 30. Platycoelia puncticollis Ohaus, 1904.................................................116 31. Platycoelia unguicularis Ohaus, 1904...............................................120 32. Platycoelia burmeisteriana Ohaus, 1917...........................................122 33. Platycoelia burmeisteri Arrow, 1899.................................................124 34. Platycoelia traceyae new species.......................................................126 35. Platycoelia wallisi Ohaus, 1904........................................................129 36. Platycoelia flavohumeralis new species.............................................131 37. Platycoelia steinheili Ohaus, 1904....................................................132 38. Platycoelia chrysotina Ohaus, 1904..................................................133 39. Platycoelia butleri new species..........................................................136 40. Platycoelia hirta Ohaus, 1904...........................................................138 41. Platycoelia altiplana new species......................................................140 42. Platycoelia confluens Ohaus, 1904....................................................141 43. Platycoelia lutescens Blanchard, 1851...............................................144 44. Platycoelia gaujoni Ohaus, 1904.......................................................149 45. Platycoelia pusilla new species.........................................................152 46. Platycoelia bordoni Martínez, 1976...................................................153 47. Platycoelia parva Kirsch, 1885..........................................................154 48. Platycoelia furva new species............................................................157 49. Platycoelia quadrilineata Burmeister, 1844.....................................159 50. Platycoelia rufosignata Ohaus, 1904................................................160 51. Platycoelia insolita new species.........................................................163 52. Platycoelia signaticollis (Burmeister, 1844).....................................166 53. Platycoelia helleri (Ohaus, 1904).......................................................169 54. Platycoelia ignota new species..........................................................171 55. Platycoelia kirschi (Ohaus, 1904)......................................................171 56. Platycoelia baessleri (Ohaus, 1904)....................................................173 57. Platycoelia aenigma new species.......................................................174 58. Platycoelia bocki (Ohaus, 1925)........................................................175 59. Platycoelia alticola (Gutiérrez, 1951)...............................................177 60. Platycoelia inca new species.............................................................179 61. Platycoelia haenkei (Gutiérrez, 1952)...............................................181 62. Platycoelia laelaps (Gutiérrez, 1951)................................................184 Acknowledgments ..........................................................................................186 Literature Cited .............................................................................................188 Appendix 1 ......................................................................................................193 Appendix 2........................................................................................................198 About the Author .............................................................................................202 iv
Page 2 and 3: Bulletin of the University of Nebra
Page 6 and 7: LIST OF FIGURES Fig. 1. Platycoelia
Page 8 and 9: Bulletin of the University of Nebra
Page 10 and 11: Members of the genus Platycoelia De
Page 12 and 13: alone to divide up the rutelines (a
Page 14 and 15: eing the most recent authority on t
Page 16 and 17: els Amongst the Great Andes of the
Page 18 and 19: tion. Zischka was a German who live
Page 20 and 21: Puncture size: large = 0.17 mm or l
Page 22 and 23: 26. Pronotal surface sparsely punct
Page 24 and 25: Fig. 11. Platycoelia alternans male
Page 26 and 27: MONOGRAPHIC REVISION OF PLATYCOELIA
Page 28 and 29: Fig. 33. Platycoelia rufosignata me
Page 30 and 31: CNCI Canadian National Collection o
Page 32 and 33: subspecies AND varieties in the sam
Page 34 and 35: PHYLOGENETIC PATTERNS AND BIOGEOGRA
Page 36 and 37: metaunguitractor plate usually with
Page 40 and 41: larvae, and Paucar-Cabrera and Smit
Page 50 and 51: Erichson (1847) did not state how m
Page 52 and 53: species]; Ohaus 1918:178 [catalog l
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MONOGRAPHIC REVISION OF PLATYCOELIA
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Platycoelia marginata and Platycoel
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tarsomere and metatarsomere 5 with
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5. Platycoelia meridensis new speci
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in apical half. Protarsomere 4 with
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7. Platycoelia prasina Erichson, 18
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written and typeface), e) “PLATYC
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mentum. Apex of mentum with triangu
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1972:302 [catalog listing as subspe
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green to yellow; parameres with the
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duced tooth. Apex of mentum with tr
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Mnisnech,” “Dr Ohaus Vidit 1903
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VARIATION. Male (n=19). Length 19.2
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“Platycoelia forcipalis cotype Oh
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males with internomedial stridulato
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tures moderate with dark coloration
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gins. Ventrally with dark brown met
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19. Platycoelia flavostriata (Latre
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20. Platycoelia nigrosternalis Ohau
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LOCALITY DATA. 66 specimens examine
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Calloodes nitidissimus Lea, 1919 (C
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