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MONOGRAPHIC REVISION OF PLATYCOELIA 169 53. Platycoelia helleri (Ohaus, 1904) (Fig. 58) Callichloris helleri Ohaus, 1904 (original combination) CATALOG. Callichloris helleri, Ohaus 1904b:333, 334, 339, 341 [original description]; Ohaus 1918:178 [catalog listing]; Blackwelder 1944:247 [checklist]; Ohaus 1952:9 [distribution]. Callichloris (Epicallichloris) helleri, Gutiérrez 1951:115, 116 [new combination, key to species of Callichloris]. Platycoelia helleri, Machatschke 1965:60 [new combination, catalog listing]; Machatschke 1972:304 [catalog listing]. TYPE SPECIMENS. Callichloris helleri Ohaus lectotype male at ZMHB labeled a) “Columb. Bogota” (typeface), b) “Typus!” (red label, typeface), c) “C. Helleri Ohaus” (red label, handwritten), d) “CALLICHLORIS HEL- LERI OHAUS M LECTOTYPE A. B. T. SMITH 2002” (red label, handwritten and typeface), e) “Platycoelia helleri (Ohaus, 1904) M Det:A.B.T.Smith 2002” (typeface). Lectotype here designated. See Methods and Materials section for a statement of taxonomic purpose. Thirteen paralectotypes were examined as follows. Three male paralectotypes at ZMHB labeled a) “Columb. Bogota” (typeface), b) “M” (typeface), c) “Callichloris Helleri Ohs. Cotype” (orange label, handwritten), d) “CALLICHLORIS HEL- LERI OHAUS M PARALECTOTYPE A. B. T. SMITH 2002” (yellow label, handwritten and typeface), e) “Platycoelia helleri (Ohaus, 1904) M Det:A.B.T.Smith 2002” (typeface). Three male paralectotypes at ZMHB labeled a) “Callanga Peru” (typeface), b) “M” (typeface), c) “Callichloris Helleri Cotype Ohs.” (orange label, handwritten), d) “CALLICHLORIS HELLERI OHAUS M PARALECTOTYPE A.B.T. SMITH 2002” (yellow label, handwritten and typeface), e) “Platycoelia helleri (Ohaus, 1904) M Det:A.B.T.Smith 2002” (typeface). One male paralectotype at SMTD labeled a) “Callauga Peru” (typeface) b) “Cotypus!” (red label, typeface), c) “76277” (green label, handwritten), d) “Callichloris Helleri Ohaus” (handwritten), e) “CALLI- CHLORIS HELLERI OHAUS M PARA- LECTOTYPE A. B. T. SMITH 2002” (yellow label, handwritten and typeface), f) “Platycoelia helleri (Ohaus, 1904) M Det:A.B.T.Smith 2002” (typeface). Two male paralectotypes at MNHN labeled a) “Colombie” (handwritten), b) “Ex.Musæo A.SALLÉ 1897” (typeface), c) “Ohaus determ. Pl. Helleri Ohaus M Cotyp.” (typeface and handwritten), d) “Dr Ohaus vidit 1903” (typeface), e) “CALLICHLORIS HELLERI OHAUS M PARALECTOTYPE A. B. T. SMITH 2002” (yellow label, handwritten and typeface), f) “Platycoelia helleri (Ohaus, 1904) M Det:A.B.T.Smith 2002” (typeface). One male paralectotype at MNHN labeled a) “Colomb.” (typeface), b) “Ex.Musæo A.SALLÉ 1897” (typeface), c) “Dr Ohaus vidit 1903” (typeface), d) “CALLICHLORIS HELLERI OHAUS M PARALECTOTYPE A.B.T. SMITH 2002” (yellow label, handwritten and typeface), e) “Platycoelia helleri (Ohaus, 1904) M Det: A.B.T.Smith 2002” (typeface). One male paralectotype (now classified as P. insolita) at MNHN labeled a) “Chancha-mayo” (handwritten), b) “Ex.Musæo H. W. Bates 1892” (typeface), c) “Ohaus determ. Pl. Helleri Ohaus M” (typeface and handwritten), d) “Dr Ohaus vidit 1903” (typeface), e) “CALLI- CHLORIS HELLERI OHAUS M PARA- LECTOTYPE A. B. T. SMITH 2002” (yellow label, handwritten and typeface), f) “PLATY- COELIA INSOLITA SMITH M PARATYPE” (yellow label, handwritten and typeface). One male paralectotype at MNHN labeled a) “Cauca Valley Colomb” (handwritten), b) “Ex.Musæo H. W. Bates 1892” (typeface), c) “Dr Ohaus vidit 1903” (typeface), d) “CALLI- CHLORIS HELLERI OHAUS M PARA- LECTOTYPE A.B.T. SMITH 2002” (yellow label, handwritten and typeface), e) “Platycoelia helleri (Ohaus, 1904) M Det:A.B.T.Smith 2002” (typeface). One male paralectotype at BMNH labeled a) “M” (typeface), b) “Co-type” (round label with yellow border, typeface), c) “Typus!” (red label, typeface), d) “Callanga, Peru 1907 - 27” (handwritten), e) “Callanga Peru” (typeface), f) “Ohaus determ. Callichloris Helleri Ohs.” (typeface and handwritten), g) “CALLICHLORIS HELLERI OHAUS M PARALECTOTYPE A.B.T. SMITH 2002” (yellow label, handwritten and typeface),
170 BULLETIN OF THE UNIVERSITYOF NEBRASKA STATE MUSEUM h) “Platycoelia helleri (Ohaus, 1904) M Det:A.B.T.Smith 2002” (typeface). Ohaus (1904b) stated that he examined about 20 male specimens in the type series. The location of the remaining (approximately seven) paratypes is unknown. Type locality: Bogotá, Colombia. DESCRIPTION. Male (n=16). Length 20.6- 22.7 mm, width 12.2-12.6 mm. Color dorsally dark green (sometimes dark yellowish-green); black frons; brown pronotal maculations; thick brown border along elytral suture, apex, lateral edge. Ventrally reddish-brown. Body ovate, convex. Head: Dorsal surface glabrous. Frons sparsely punctate (base, lateral edge) to densely punctate (medioapically), clypeus rugose, punctures moderate. Frons weakly depressed medioapically. Frontoclypeal suture complete, weakly bisinuate. Clypeus rectangular, apex slightly reflexed. Eyes separated by approximately 4.5 transverse eye-widths. Labrum rugose with setose punctures, setae tawny. Apex of labrum with minute, triangular, medial tooth, apex of tooth well separated from apex of mentum. Mandibular scissorial region with 1 tooth, molar region with weak lamellae. Maxilla with 3 apical teeth. Apex of mentum with medial notch. Antenna 9-segmented; club approximately equal to segments 2-6, approximately equal to clypeal length. Pronotum: Surface glabrous, sparsely punctate, with small and moderate punctures. Marginal bead present apically (except medially) and laterally, absent basally. Elytron: Surface glabrous; longitudinal striae weakly impressed, weakly punctate; intervals impunctate. Suture apically rounded. Pygidium: Width approximately 1.7 times length medially. Surface smooth; disc flat, impunctate (except moderately punctate near apex); punctures moderately large, setose; setae long, tawny. Venter: Thorax densely setose; setae long, tawny to golden-brown. Mesothoracic process barely produced, projecting anteriorly to subapex of mesocoxa. Abdomen sparsely setose laterally; setae long, tawny. Apical spiracles not extruding. Legs: Protibia with 3 teeth in apical half; first and second teeth subequal in size; third tooth small, often worn and obsolete, removed slightly from apical teeth. Mesotibia and metatibia widest apically. Protarsomeres 2-4 wider than long, cup-shaped. Protarsomere 3-4 with internoapical stridulatory ridge. Protarsomere 5 with internomedial stridulatory tooth. Mesotarsomere and metatarsomere 5 without internal tooth. Unguitractor plate cylindrical, with 1 apical and 1 subapical seta. Modified protarsal claw with length approximately equal to protarsomere 5, thickened and elongated when compared with other claw, dorsoventrally flattened, apex unevenly bifurcate. Modified mesotarsal and metatarsal claws elongated with ventral tooth, not thickened. Male Genitalia: Phallobase 1.6 times longer than length of parameres. Parameres with apex rounded. Female: Unknown. DIAGNOSIS. This species is distinguished from all other species in the genus Platycoelia by the following combination of characters: frons dark in color; frontoclypeal suture complete; apex of the labrum with a reduced, triangular tooth not overlapping the mentum; mentum with a medial notch; antenna 9-segmented; pronotum with weak maculations; elytron without distinct, yellow, longitudinal lines; elytral apices not capable of completely covering the pygidium; elytron with a thick, black suture line; apex of the elytral suture without an acute spine; mesothoracic process nub to weakly projecting apically; apical abdominal spiracle not protuberant; mesotarsomere 5 and metatarsomere 5 without an internal tooth; parameres with the apex rounded, not expanded. DISTRIBUTION (Fig. 58). If the data labels are to be believed, this species occurs in highly disjunct areas of Andean Colombia and Perú. More specimens with reliable data are needed to determine the distribution of this species. LOCALITY DATA. 16 specimens examined from BMNH, MNHN, SMTD, ZMHB. COLOMBIA (9). ANTIOQUIA (1): Medellín. DISTRITO CAPITAL (4): Bogotá. NO DATA (4) PERÚ (7). LIMA (5): Callanga. NO DATA (2).
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Bulletin of the University of Nebra
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C O N T E N T S Abstract...........
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LIST OF FIGURES Fig. 1. Platycoelia
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Bulletin of the University of Nebra
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Members of the genus Platycoelia De
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alone to divide up the rutelines (a
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eing the most recent authority on t
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els Amongst the Great Andes of the
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tion. Zischka was a German who live
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Puncture size: large = 0.17 mm or l
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26. Pronotal surface sparsely punct
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Fig. 11. Platycoelia alternans male
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MONOGRAPHIC REVISION OF PLATYCOELIA
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Fig. 33. Platycoelia rufosignata me
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CNCI Canadian National Collection o
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subspecies AND varieties in the sam
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PHYLOGENETIC PATTERNS AND BIOGEOGRA
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metaunguitractor plate usually with
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larvae, and Paucar-Cabrera and Smit
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Erichson (1847) did not state how m
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species]; Ohaus 1918:178 [catalog l
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Platycoelia marginata and Platycoel
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tarsomere and metatarsomere 5 with
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5. Platycoelia meridensis new speci
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in apical half. Protarsomere 4 with
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7. Platycoelia prasina Erichson, 18
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written and typeface), e) “PLATYC
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mentum. Apex of mentum with triangu
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1972:302 [catalog listing as subspe
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green to yellow; parameres with the
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duced tooth. Apex of mentum with tr
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Mnisnech,” “Dr Ohaus Vidit 1903
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VARIATION. Male (n=19). Length 19.2
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“Platycoelia forcipalis cotype Oh
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males with internomedial stridulato
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tures moderate with dark coloration
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gins. Ventrally with dark brown met
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19. Platycoelia flavostriata (Latre
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20. Platycoelia nigrosternalis Ohau
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LOCALITY DATA. 66 specimens examine
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Page 126 and 127: MONOGRAPHIC REVISION OF PLATYCOELIA
Page 206 and 207: Calloodes nitidissimus Lea, 1919 (C
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