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MONOGRAPHIC REVISION OF PLATYCOELIA 145 be set aside on the grounds that they do not comply with the changed provisions of the fourth edition. The Commission should be asked to validate the names or acts.” A request to the Commission to uphold this lectotype designation has been published (Smith 2002b). One male paralectotype at MNHN labeled a) “Machachi, Ecuador. 9-10,000 feet. Ed Whymper” (typeface) on upperside and “1368171 K” (handwritten) on lowerside, b) “Leucopelaea albescens Bates” (handwritten), c) “Ex-Musæo H.W.BATES 1892” (typeface), d) “Dr Ohaus Vidit 1903.” (typeface), e) “LEU- COPELAEA ALBESCENS BATES 1891 DET:A.B.T.SMITH 2001 PARALECTOTYPE” (yellow label, handwritten and typeface), f) “PLATYCOELIA LUTESCENS BLAN- CHARD M Det:A.B.T.Smith 2001” (handwritten and typeface). One male paralectotype at MNHN labeled a) “Cotopaxi 12000 ft” (handwritten), b) “Leucopelaea albescens Bates” (handwritten), c) “Ex-Musæo H.W.BATES 1892” (typeface), d) “Dr Ohaus Vidit 1903.” (typeface), e) “LEUCOPELAEA ALBESCENS BATES 1891 DET:A.B.T.SMITH 2001 PARALECTOTYPE” (yellow label, handwritten and typeface), f) “PLATYCOELIA LUTESCENS BLANCHARD M Det: A.B.T. Smith 2001” (handwritten and typeface). Four male paralectotypes at MNHN labeled a) “Cotopaxi 12000 ft” (handwritten), b) “Ex- Musæo H.W.BATES 1892” (typeface), c) “Dr Ohaus Vidit 1903.” (typeface), d) “LEUCO- PELAEA ALBESCENS BATES 1891 DET: A.B.T.SMITH 2001 PARALECTOTYPE” (yellow label, handwritten and typeface), e) “PLATYCOELIA LUTESCENS BLANCHARD M Det:A.B.T.Smith 2001” (handwritten and typeface). One male paralectotype at ZMHB labeled a) “Cotopaxi 12000” (handwritten), b) “Leucopelaea albescens Cotype M Bts.” (orange label, handwritten), c) “LEUCOPELAEA ALBESCENS BATES 1891 DET: A.B.T. SMITH 2001 PARALECTOTYPE” (yellow label, handwritten and typeface), d) “PLATY- COELIA LUTESCENS BLANCHARD M Det:A.B.T.Smith 2001” (handwritten and typeface). One male paralectotype at ZMHB labeled a) “Machachi Ecuador” (typeface), b) “M” (typeface), c) “Leucopelaea albescens H. Bates” (handwritten), d) “LEUCOPELAEA ALBESCENS BATES 1891 DET: A.B.T. SMITH 2001 PARALECTOTYPE” (yellow label, handwritten and typeface), e) “PLATY- COELIA LUTESCENS BLANCHARD M Det:A.B.T.Smith 2001” (handwritten and typeface). Bates (1891b) stated that there were 13 specimens in the type series. The location of the remaining 4 paralectotypes is unknown. Placed in synonymy with P. lutescens by Smith and Paucar-Cabrera (2000). Type locality: the plain of Limpiopungo near Volcán Cotopaxi, Ecuador (Whymper 1892). Leucopelaea baronis Ohaus lectotype male at ZMHB labeled a) “Ecuador Baron” (typeface), b) “Leucopelaea Baronis Type Ohs.” (orange label, handwritten), c) “LEU- COPELAEA BARONIS OHAUS M LECTO- TYPE A. B. T. SMITH 2002” (red label, handwritten and typeface), d) “Zool. Mus. Berlin” (typeface), e) “Platycoelia LUTESCENS BLANCHARD M Det: A.B.T. Smith 1999” (typeface and handwritten). Lectotype here designated. See Methods and Materials section for a statement of taxonomic purpose. Two male paralectotypes at ZMHB labeled a) “Ecuador Baron” (typeface), b) “Leucopelaea Baronis Cotype Ohs.” (orange label, handwritten), c) “LEUCOPELAEA BARONIS OHAUS M PARALECTOTYPE A. B. T. SMITH 2002” (yellow label, handwritten and typeface), d) “Platycoelia LUTESCENS BLANCHARD M Det:A.B.T.Smith 1999” (typeface and handwritten). Two female paralectotypes at ZMHB labeled a) “Ecuador Baron” (typeface), b) “Leucopelaea Baronis Cotype Ohs.” (orange label, handwritten), c) “LEUCOPELAEA BARONIS OHAUS F PARALECTOTYPE A. B. T. SMITH 2002” (yellow label, handwritten and typeface), d) “Platycoelia LUTESCENS BLANCHARD F Det:A.B.T.Smith 1999” (typeface and handwritten). Three male paralectotypes at ZMHB labeled a) “Ecuador Baron” (typeface), b) “Leucopelaea Baronis Cotype Ohs.” (orange label, handwritten), c) “LEUCOPELAEA BARONIS OHAUS M PARALECTOTYPE A.B.T. SMITH 2002” (yellow label, handwritten and typeface), d) “PLATYCOELIA LUTESCENS BLAN-
146 BULLETIN OF THE UNIVERSITYOF NEBRASKA STATE MUSEUM CHARD M Det:A.B.T.Smith 2001” (handwritten and typeface). Two female paralectotypes at ZMHB labeled a) “Ecuador Baron” (typeface), b) “Leucopelaea Baronis Cotype Ohs.” (orange label, handwritten), c) “LEUCOPEL- AEA BARONIS OHAUS F PARALECTO- TYPE A.B.T. SMITH 2002” (yellow label, handwritten and typeface), d) “PLATYCOE- LIA LUTESCENS BLANCHARD F Det: A.B.T. Smith 2001” (handwritten and typeface). Smith and Paucar-Cabrera (2000) erroneously referred to the type specimens of Leucopelaea baronis as a holotype and paratypes (see discussion of Ohaus’ type in Methods and Materials section). One specimen labeled “Ecuador E. Deville” also had an orange “Leucopelaea Baronis Cotype Ohs.” label but the original description mentioned only specimens collected by Baron (Ohaus 1905) so this specimen is not considered to be in the type series. Placed in synonymy with P. lutescens by Smith and Paucar-Cabrera (2000). Type locality: Ecuador. DESCRIPTION. Male (n=226). Length 16.1- 24.3 mm, width 10.2-13.6 mm (Fig. 70). Color dorsally and ventrally cream-colored when alive (often turning tan or testaceous when dead). Body ovate, convex. Head: Dorsal surface glabrous. Frons impunctate or sparsely punctate (base) to moderately punctate (apex), punctures moderate. Clypeus densely punctate (base) to rugopunctate (apex), punctures moderate. Frons not depressed. Frontoclypeal suture complete, weakly bisinuate. Clypeal apex rounded, moderately reflexed. Eyes separated by approximately 5.5 transverse eye-widths. Labrum rugose, setose; setae tawny. Apex of labrum with minute, triangular, medial tooth, apex of tooth well separated from apex of mentum. Mandibular scissorial region with weak apical tooth, weak medial nub; molar region without lamellae. Maxilla with 2-3, apical teeth. Apex of mentum with medial notch. Antenna 10-segmented (sometimes appearing 9-segmented); club slightly shorter than other segments combined, slightly longer than clypeal length. Pronotum: Surface glabrous, sparsely punctate, with moderate punctures. Marginal bead weak laterally, absent apically and basally. Elytron: Surface glabrous, with weakly impressed longitudinal striae; intervals impunctate. Suture apically rounded to angular. Pygidium: Width approximately 1.7 times length. Surface weakly convex, punctate; punctures small, sparse, setose in apical half; setae long, tawny. Venter: Thorax moderately to densely setose; setae long, tawny. Mesothoracic process small, projecting anteriorly to subapex of mesocoxa, or reduced nub. Abdominal sternites laterally moderately setose, medially sparsely setose; setae long, reddish. Apical 2 spiracles with margins strongly extruding, cylindrical. Legs: Protibia with 3 teeth in apical half; first and second teeth subequal in size in size, third tooth small, often worn and obsolete, removed slightly from apical teeth. Mesotibia and metatibia widest medially. Protarsomeres 2-4 wider than long, cup-shaped. Protarsomere 3-4 without internoapical stridulatory ridge. Protarsomere 5 with internomedial tooth or swelling. Mesotarsomere and metatarsomere 5 without internal tooth. Unguitractor plate cylindrical, with 1 apical, 1 subapical seta. Modified protarsal claw with length approximately equal to protarsomere 5, thickened and elongate when compared with other claw, dorsoventrally flattened, apex unevenly bifurcate. Modified mesotarsal and metatarsal claws elongated with ventral tooth, not thickened. Male Genitalia: Phallobase 1.4 times longer than length of parameres. Parameres with apex rounded. Female (n=35). Length 17.5-30.3 mm, width 10.7-16.8 mm. As male except in the following respects. Head: Antennal club approximately equal to segments 2-7. Legs: Third protibial tooth broadly acute, rarely worn down and obsolete. Protarsomere 5 without internal tooth. Modified protarsal claw with ventral tooth, not thickened. LARVAE. Platycoelia lutescens larvae were described by Paucar-Cabrera and Smith (2002). The following diagnostic characters were given: epipharynx with laeotorma longer than dexiotorma, expanded towards chaetoparia; mandible with two scissorial teeth; abdominal spiracles elongate.
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Bulletin of the University of Nebra
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C O N T E N T S Abstract...........
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LIST OF FIGURES Fig. 1. Platycoelia
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Bulletin of the University of Nebra
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Members of the genus Platycoelia De
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alone to divide up the rutelines (a
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eing the most recent authority on t
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els Amongst the Great Andes of the
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tion. Zischka was a German who live
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Puncture size: large = 0.17 mm or l
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26. Pronotal surface sparsely punct
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Fig. 11. Platycoelia alternans male
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MONOGRAPHIC REVISION OF PLATYCOELIA
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Fig. 33. Platycoelia rufosignata me
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CNCI Canadian National Collection o
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subspecies AND varieties in the sam
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PHYLOGENETIC PATTERNS AND BIOGEOGRA
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metaunguitractor plate usually with
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larvae, and Paucar-Cabrera and Smit
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Erichson (1847) did not state how m
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species]; Ohaus 1918:178 [catalog l
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Platycoelia marginata and Platycoel
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tarsomere and metatarsomere 5 with
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5. Platycoelia meridensis new speci
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in apical half. Protarsomere 4 with
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7. Platycoelia prasina Erichson, 18
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written and typeface), e) “PLATYC
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mentum. Apex of mentum with triangu
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1972:302 [catalog listing as subspe
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green to yellow; parameres with the
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duced tooth. Apex of mentum with tr
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Mnisnech,” “Dr Ohaus Vidit 1903
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VARIATION. Male (n=19). Length 19.2
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“Platycoelia forcipalis cotype Oh
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males with internomedial stridulato
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tures moderate with dark coloration
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gins. Ventrally with dark brown met
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19. Platycoelia flavostriata (Latre
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Page 102 and 103: 20. Platycoelia nigrosternalis Ohau
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Calloodes nitidissimus Lea, 1919 (C
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