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eing the most recent authority on the subject.<br />

The publication dates for the fascicles in<br />

which Platycoelia was cataloged were listed in<br />

Madge (1988) and confirmed by the journal<br />

Bibliographie de la France which announces<br />

the release <strong>of</strong> new publications in that country.<br />

Platycoelia Dejean, 1833 was published no<br />

later than 27 July 1833 according to the<br />

Bibliographie de la France (36[30]: 459). Although<br />

Dejean (1833) provided no description<br />

for the new generic name, Platycoelia was<br />

made available in that publication under the<br />

ICZN rules (Article 12.2.5). The placement <strong>of</strong><br />

an available specific name in combination<br />

with a new generic name makes the generic<br />

name available by indication (International<br />

Commission on Zoological Nomenclature<br />

1999). Ohaus (1918) and Machatschke (1965)<br />

incorrectly (according to the current rules <strong>of</strong><br />

nomenclature) cited the author and date <strong>of</strong><br />

Platycoelia as Burmeister, 1844.<br />

Burmeister (1844) was the first author to<br />

attempt to classify Platycoelia in the con<strong>text</strong><br />

<strong>of</strong> other groups <strong>of</strong> scarab beetles. He was the<br />

first to provide a description <strong>of</strong> the genus and<br />

he placed the group in the “Anoplognathidae,”<br />

which is the modern tribe Anoplognathini consisting<br />

<strong>of</strong> Australian and Neotropical genera.<br />

Burmeister also erected the family group<br />

name Platycoeliina Burmeister (originally as<br />

Platycoeliidae) containing Platycoelia and<br />

Phalangogonia, and he discussed similarities<br />

with Anoplognathus Leach and Repsimus<br />

MacLeay (two Australian genera). Three new<br />

species <strong>of</strong> Platycoelia were also described in<br />

this paper: P. valida Burmeister, P. quadrilineata<br />

Burmeister, and P. marginata Burmeister.<br />

Burmeister also redescribed P.<br />

flavostriata, but used specimens <strong>of</strong> another<br />

species for the description. The species<br />

Burmeister described as P. flavostriata was<br />

later recognized as different by Arrow (1899)<br />

and Ohaus (1904b) and given the names P.<br />

burmeisteri Arrow and P. tschudii Ohaus respectively<br />

(with the Arrow name being the<br />

valid one). Burmeister (1844) also described<br />

P. signaticollis (Burmeister) in the new genus<br />

Callichloris, which he placed under the family<br />

group name Brachysternidae (with taxa<br />

currently in the subtribes Brachysternina<br />

and Schizognathina). Callichloris was also<br />

MONOGRAPHIC REVISION OF PLATYCOELIA 7<br />

listed by Dejean (1833, 1836), but was not<br />

made available in those publications because<br />

no available species-group names were listed<br />

under that generic name.<br />

Erichson (1847) described three new species<br />

<strong>of</strong> Platycoelia from Perú: P. pomacea<br />

Erichson, P. alternans Erichson, and P.<br />

prasina Erichson. Accession records at ZMHB<br />

indicate that the type series <strong>of</strong> P. alternans<br />

was collected by Rudolph A. Philippi and the<br />

type series <strong>of</strong> P. pomacea and P. prasina were<br />

collected by José Pavón (Pavón specimens arrived<br />

at ZMHB via the H<strong>of</strong>fmansegg collection).<br />

Blanchard (1851) later described three<br />

more species: P. boliviensis Blanchard (herein<br />

a junior synonym <strong>of</strong> P. pomacea); P. olivacea<br />

Blanchard (later placed in synonymy with P.<br />

marginata); and P. lutescens Blanchard.<br />

Burmeister (1855) later added one additional<br />

species, P. laevis Burmeister (later placed in<br />

synonymy with P. marginata). In the same<br />

year, Guérin-Méneville (1855) described another<br />

species, P. scutellata Guérin-Méneville<br />

(herein a junior synonym <strong>of</strong> P. marginata).<br />

Kirsch (1871, 1885) then added two species:<br />

P. nervosa Kirsch and P. parva Kirsch. Bates<br />

(1888, 1891b) described two new species: P.<br />

humeralis Bates and P. nigricauda Bates<br />

(herein a junior synonym <strong>of</strong> P. parva). Bates<br />

(1891b) also described the new genus Leucopelaea<br />

Bates (later placed in synonymy with<br />

Platycoelia) to accommodate the new species<br />

L. albescens Bates (later placed in synonymy<br />

with P. lutescens).<br />

Before the 20th century, there was no<br />

comprehensive work done on the genus<br />

Platycoelia. Freidrich Ohaus, who has been<br />

called the “Father <strong>of</strong> Rutelinae” (Jameson<br />

1998), changed all that with the first review<br />

<strong>of</strong> the genus in 1904. Ohaus (1904b) redescribed<br />

all the existing species and described<br />

the following 23 new species group<br />

names <strong>of</strong> Platycoelia: P. inflata Ohaus, P.<br />

inflata tucumana Ohaus (herein a junior synonym<br />

<strong>of</strong> P. inflata), P. limbata Ohaus (herein<br />

a junior synonym <strong>of</strong> P. prasina), P. steinheili<br />

Ohaus, P. forcipalis Ohaus, P. occidentalis<br />

Ohaus, P. pomacea brasiliensis Ohaus (herein<br />

a junior synonym <strong>of</strong> P. pomacea), P.<br />

abdominalis Ohaus, P. mesosternalis Ohaus,<br />

P. variolosa Ohaus, P. nigrosternalis Ohaus,

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